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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Family Ranidae Batsch, 1796 
Genus Amnirana Dubois, 1992 
Amnirana albolabris (Hallowell, 1856) White-Lipped Frog 
Rana albolabris Hallowell 1856:153. Syntypes: ANSP [4 specimens], lost fide J.L. Ferret in Frost (1985:479) 
(collector P. du Chaillu). Type locality: “W. Africa” (Hallowell 1856:153), restricted to “Gabon” by Ferret 
(1977:843). 
Limnodytes albolabris: Peters (1877a:618). 
Rana albolabris: Bocage (1895a: 162). 
Hylarana albolabris albolaris: Laurent (1950a: 14, 1954a:74), Loveridge (1957:338). 
Hylarana albolabris: Ferret (1977:843), Ruas (1996:24). 
Amnirana albolabris: Channing and Howell (2006:256), Channing et al. (2012:360), Frost (2016), Jongsma 
et al. (2018:275). 
Rana (Amnirana) albolabris: Fretey et al. (2011:43). 
[-5.10000, 12.10000] (Peters 1877a:618; Boca¬ 
ge 1895a; 162). Uige: “Uige, University Kimpa 
Vita campus” [-8.607967, 20.968804] (Jongs¬ 
ma et al. 2018:275); “Serra do Pingano” 
[-7.68528, 14.92956] (Jongsma et al. 
2018:275); “Negage” [-7.7625, 15.285] 
(Jongsma et al. 2018:275); “Uige fish farm, 
Kibokolo” [-6.274417, 15.067472] (Jongsma 
et al. 2018:275). Lunda Norte: “Dundo” 
[-7.36667, 20.83333] (Laurent 1950a: 14, 
1954a:74; Ruas 1996:24); “Muita (Luembe E)” 
[-7.80000, 21.45000] (Laurent 1950a:14; Ruas 
1996:24); “rive de la Tshihumbe, 40 km a Lest 
de Dundo (Rio Chiumbe)” [-8.01667, 
19.31667] (Laurent 1950a: 14; Ruas 1996:24); 
“Tshinguvu, galerie forestiere de la Tshikapa, 
50 km au sub-ouest de Dundo (Chingufo)” [-8.01667, 19.31667] (Laurent 1950a: 14; Ruas 
1996:24); “Sombo” [-8.68333, 20.95000] (Laurent 1954a:74; Ruas 1996:24). 
Taxonomic and distributional notes: Ferret (1977) restricted the distribution of this species 
to the forested area of Cameroon and Congo (Channing and Howell 2006). Laurent (1964a) was 
uncertain whether his Angolan material corresponded to A. albolabris or instead to A. lemairei 
(Laurent 1950a, 1954a). Jongsma et al. (2018) included Angolan populations in their molecular 
phylogenetic analyses and found that these fell within a large complex of likely cryptic species all 
currently referred to as A. albolabris. 
Map 104. Distribution of Amnirana albolabris in 
Angola. 
Global conservation status (lUCN): Least Concern. 
Global distribution: The wide-ranging forest species found across western and Central Africa 
likely represents multiple species, each with more restricted distributions. 
Ocurrences in Angola (Map 104): The 
species occurs in northern Angola and in the 
Cabinda enclave. Cabinda: “Chinchoxo” 
Amnirana darlingi (Boulenger, 1902) Darling’s White-Lipped Frog 
Rana darlingi Boulenger 1902:15. Syntypes: BMNH 1947.2.2.67-68 (eolleetor J. ffolliott Darling), formerly 
