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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
highlands” (Frade 1963:254); “areas of forest and savanna in the north and northeast of Angola” 
(Cei 1977:16). 
Taxonomic and distributional notes: Laurent (1964a) synonymized Rana albolabris adis- 
cifera with Rana darlingi. 
Amnirana lemairei (de Witte, 1921) Lemaire’s White-Lipped Frog 
Rana Lemairei de Witte 1921:1, pi. 1, fig. 1 -4. Holotype: MRAC 172 (eolleetor Mission Lemaire). Type loeal- 
ity: “Lofoi (Katanga)” (de Witte 1921:3), Demoeratie Republie of Congo. 
Hylarana albolabris lemairei: Laurent (1964a: 133), Cei (1977:16) 
Hylarana lemairei: Poynton and Broadley (1985b: 140), Ruas (1996:24, 2002:143), Channing et al. 
(2012:365). 
Amnirana lemairei: Channing (2001:269), Frost (2016). 
Rana {Amnirana) lemairei: Fretey et al. (2011:43). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from northeastern Angola, the southern Demo¬ 
cratic Republic of Congo, and western Zambia. 
Ocurrences in Angola (Map 106): The 
species occurs in northeastern Angola. Lunda 
Norte: “rives du Luachimo, Dundo (Luachi- 
mo)” [-7.38333, 20.85000] (Laurent 
1964a: 133; Ruas 1996:24); “Riv. Chitato, 
Dundo (Rio Chitado)” (Laurent 1964a: 133; 
Ruas 1996:24). Lunda Sul: “Alto Cuilo, dans 
I’eau, tres basse, des sources d’un ruisseau 
(Alto Cuilo)” [-10.01667, 19.55000] (Laurent 
1964a: 133; Ruas 1996:24); “Chutes du Cuan- 
go-Muque, Alto Chicapa (Rio Cuango-Muque, 
quedas)” [-10.76667, 19.20000] (Laurent 
1964a: 133; Ruas 1996:24); “sources de la 
Tchimboma, affl. gauche du Cuango-Muque, 
Alto Chicapa (Rio Chimboma, nascente)” 
[-10.93333, 19.18333] (Laurent 1964a:133; 
Ruas 1996:24). Mexico: “Lumeje I, galerie 
forestiere isolee a cote de la route, pres de I’en¬ 
Map 106. Distribution of Amnirana lemairei in Angola. 
tree de la reserve de chasse de Cameia (Rio Lumege)” [-11.58333, 21.00000] (Laurent 1964a: 133; 
Ruas 1996:24); “Lago Cameia” [-11.71667, 20.80000] (Ruas 1996:24). Undetermined Locality: 
“areas of forest and savanna in the north and northeast of Angola” (Cei 1977:16). 
Taxonomic and distributional notes: This taxon was discussed by Schmidt and Inger (1959) 
as a subspecies of Rana albolabris (Hallowell, 1856). Laurent (1964a) identified some individuals 
as Rana albolabris lemairei, and Perret (1977) later recognized this as a full species, although he 
expressed some doubts about it occurence in Angola. 
Amnirana lepus (Andersson, 1903) Andersson’s White-Lipped Frog 
Chiromantis lepus Andersson 1903: 142. Holotype: NHR ? (eolleetor Y. Sjostedt). Type loeality: “Kamemn,” 
[=Cameroon]. 
Amnirana lepus: Jongsma et al. (2018:275). 
