MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Pelomedusa subrufa: Iverson (1986:236, 1992:52), Boycott and Bourquin (2008:007.1), Turtle Taxonomy 
Working Group (2014:435), Petzold et al. (2014:542), Ceriaco et al. (2016a:54), Conradie et al. (2016:26). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is distributed from the subtropical savannas and semi-desert 
regions in Angola, Namibia, Botswana, Zimbabwe, Zambia, Malawi, the southeastern Democratic 
Republic of Congo, parts of Mozambique, and portions of northeastern South Africa. It has also 
been introduced to Madagascar. 
Ocurrences in Angola (Map 110): This 
species has been confirmed from numerous 
localities across southern and central Angola. 
Malanje: “Duque de Braganqa” [-9.10000, 
15.95000] (Bocage 1895a:5; Monard 
1937b: 146; Loveridge 1941b:479). Bie: 
“Gauca” [-11.18333, 17.45000] (Schmidt 
1933:3; Loveridge 1941b;479); “Chitau” 
[-11.43333, 17.15000] (Schmidt 1933:3; Love¬ 
ridge 1941b;479). Benguela: “Quissange” 
[-12.43333, 14.05000] (Bocage 1895a:5; 
Monard 1937b: 146; Loveridge 1941b: 
479); “Catumbella-Miindung, Benguella” 
[-12.33333, 13.55000] (Mertens 1926:152). 
Huila: “Kalukembe” [-13.78333, 14.68333] 
(Monard 1937b:147; Loveridge 1941b:479); 
“Quillengues” [-14.06667, 15.08333] (Bocage 
1895a:5; 1937b:146; Loveridge 1941b:479); 
Map 110. Distribution of Pelomedusa subrufa in Angola. 
“sur les hordes du Kuvangu, au sud de Vila-da-Ponte (Kuvangu river S. Vila da Ponte)” [-14.46667, 
16.30000] (Monard 1937b: 147; Loveridge 1941b:479); “Huila District” [-15.24179, 14.52210] 
(Petzold et al. 2014, supplemental information); “Kahuihui” [Undetermined Locality] (Monard 
1937b: 147; Loveridge 1941b:479); “Kalundunga” [Undetermined Locality] (Monard 1937b: 147; 
Loveridge 1941b:479). Namibe: “Mucungu” [-14.78333, 12.48333] (Schmidt 1933:3; Loveridge 
194bl:479; Ceriaco et al. 2016a:54); “Maconjo” [-15.01667, 13.20000] (Bocage 1895a:5; Monard 
1937b:146; Loveridge 1941b:479; Ceriaco et al. 2016a:54); “Capangombe” [-15.10000, 13.15000] 
(Bocage 1887b:202; 1895a:5; Monard 1937b:146; Loveridge 1941b:479; Ceriaco et al. 2016a:54); 
“Mossamedes” [-15.20000, 12.15000] (Bocage 1887b:202; 1895:5; Loveridge 1941b:479; Ceria¬ 
co et al. 2016b:54). Cunene: “Humbe” [-16.68333, 14.90000] (Bocage 1895a:5; 1937b:146; 
Loveridge 1941b:479). Cuando Cubango: “23 km E Jamba” [-17.48690,22.98790] (Petzold et al. 
2014, supplemental information); “Cuando basin (40)” [-17.45786, 22.91191] (Conradie et al. 
2016:9-10, 26). 
Taxonomic and distributional notes: This species was first described by Lacepede (1788) as 
Testudo subrufa, but following the ICZN ruling that this was an unavailable non-binominal work 
(ICZN 2005, Opinion 2104, Case 3226), authorship has been attributed to Bonnaterre (1789). 
Mertens (1937b) and Loveridge (1941b) demonstrated that T. subrufa pre-dated T. galeata, until 
then widely employed as the name for the helmeted terrapin. As the collector named by Lacepede, 
Pierre Sonnerat, was known to have collected in the southwestern Cape region of South Africa, 
Mertens (1937b) restricted the type locality to this area. However, Bour (1982) presented evidence 
that the type was probably actually collected by Philibert Commerson in Madagascar in 1770 
(see review in Fritz et al. 2014). Loveridge (1941b) cited several species and subspecies for 
