MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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as P. castaneus (Schweigger, 1812). Raw (1978) elevated P. rhodesianus Hewitt, 1927 to specific 
rank, an action subsequently confirmed by Broadley (1981a) and accepted by most subsequent 
authors. Iverson (1987, 1992) presented a point distribution map for the species, although he did 
not provide information about the localities depicted. Broadley and Boycott (2008) also provided 
a distribution map of P. rhodesianus based on museum and literature records. 
Family Testudinidae Batsch, 1788 
Genus Kinixys Bell, 1827 
Kinixys belliana and Kinixys spekii 
The precise delineation of these two species of Kinixys in Angola remains unclear and genet¬ 
ic data will be necessary to establish species boundaries (U. Fritz, pers. comm.). To date only 
K. belliana has been genetically verified in Angola but it is highly likely, given the distribution of 
K. spekii elsewhere, that it is also present in the country (see notes below). Collectively the species 
are widespread, although absent from desert and forest habitats. 
Ocurrences in Angola (Map 116): The 
following records combine both K. belliana 
and K. spekii (see K. spekii account below for 
the specific reference to its records in the cher- 
sonyms list). Lunda Norte: “Muita (Luembe 
E)” [-7.80000, 21.45000] (Laurent 1950a;13; 
Loveridge and Williams 1957:396). Lunda 
Sul: “Alto Chicapa” [-10.88333, 19.23333] 
(Laurent 1964a:25); “Lunda” [-10.96667, 
20.06667] (Monard 1937b; 146; Loveridge and 
Williams 1957:396). Moxico: “environs du lac 
Calundo” [-11.80000, 20.86667] (Laurent 
1964a:25). Malanje: “Duque de Braganga” 
[-9.10000, 15.95000] (Bocage 1866a:40, 
1895a:2; Loveridge and Williams 1957:396); 
“Muata-Yamv, Lui River” [-9.41667, 
17.33333] (Loveridge and Williams 1957:396); 
“Capanda” [-9.72841,15.34585] (Ceriaco et al. 
2014b:670); “Cangandala National Park” 
[-9.84606, 16.72233] (Ceriaco et al. 2016b:53). Bie: “Gauca” [-11.18333, 17.45000] (Laurent 
1964a:25; Loveridge and Williams 1957:396); “Chitau” [-11.43333, 17.15000] (Schmidt 1933:4; 
Loveridge and Williams 1957:396); “Cuemba” [-12.15000, 18.08333] (Kindler et al. 2012, 
supporting information:4); “Kuito” [-12.383599, 16.928029] (Kindler et al. 2012, supporting infor- 
mation;4); “Cubando basin (13)” [-13.28061,16.74722] (Conradie et al. 2016a:8-9,27). Huambo: 
“Galanga” [-12.06667, 15.15000] (Bocage 1895a:2; Loveridge and Williams 1957:396). 
Benguela: “Quissange” [-12.43333, 14.05000] (Bocage 1887b:209, 1895a;2; Loveridge and 
Williams 1957:396); “Benguella” [-12.58333, 13.41667] (Hellmich 1957a:32); “Ebanga” 
[-12.73333, 14.73333] (Monard 1937b; 146; Loveridge and Williams 1957:396); “Cubal” 
[-13.03333, 14.25000] (Mertens 1937a:5, 1938a:430; Loveridge and Williams 1957:396). Hufla: 
“Caconda” [-13.73333, 15.06667] (Bocage 1895a;2; Loveridge and Williams 1957:395); “Kuvan- 
gu” [-14.46667, 16.30000] (Monard 1937b:146); “Osi” [-15.08333, 15.41667] (Monard 
1937b:146; Loveridge and Williams 1957:396). Cunene: “riv. Mbale” [-15.16667, 16.75000] 
Map 116. Distribution of Kinixys belliana and Kinixys 
spekii in Angola. 
