MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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ed it to the Oxford University Museum in 1862. However, a recent catalogue of the types there 
reveals that the type of S. pardalis is not present (Nowak-Kemp and Fritz 2010). Stigmochelys 
pardalis (Bell, 1828) was previously included in the genus Testudo and more recently, in the genus 
Geochelone (Loveridge and Williams 1957; Iverson 1992). Based on data from mitochondrial and 
nuclear genes, Le et al. (2006:524, 528) indicated a sister relationship between S. pardalis and 
Psammobates and subsumed it within the latter genus. However, based on morphological differ¬ 
ences, Fritz and Bininda-Emonds (2007) placed it in a monotypic Stigmochelys, which has been 
accepted by subsequent authors. Loveridge and Williams (1975) recognized two subspecies for 
Geochelone (= Stigmichelis) pardalis (Bell, 1828) G. p. pardalis and G. p. babcocki, also distin¬ 
guished by Le et al. (2006) and Fritz and Havas (2007). Iverson (1992) formerly questioned the 
recognition of the two subspecies and recently Fritz et al. (2010) concluded that there is no ration¬ 
ale for recognizing subspecies within S. pardalis. Broadley (1989b) and Iverson (1986, 1992) pre¬ 
sented distribution maps for the species with some new Angolan localities but without mentioning 
the localities by name. 
Family Trionychidae Fitzinger, 1826 
Genus Cycloderma Peters, 1854 
Cycloderma aubryi (Dumeril, 1856) Aubry’s Softshell Turtle 
Cryptopodus Aubryi Dumeril 1856:374, pi. 20. Holotype, MNHN 8006 (collector C.E. Aubry-Lecomte). Type 
locality: “Gabon.” 
Cycloderma Aubryi: Peters (1877a:611), Bocage (1895a:8). 
Cycloderma aubryi: Loveridge and Wiliams (1957:453), Frade (1963:252), Iverson (1992:295), Gramentz 
(2008:117), Turtle Taxonomy Working Group (2014:411). 
Global conservation status (lUCN): Vul¬ 
nerable. 
Global distribution: Central African 
species known from the Congo Basin from 
Gabon to the Democratic Republic of Congo 
and Cabinda in Angola. 
Ocurrences in Angola (Map 119): This 
species occurs in the Cabinda enclave. Cabin¬ 
da: “Chinchoxo” [-5.10000, 12.10000] (Peters 
1877a:611; Bocage 1895a:7; Loveridge and 
Williams 1957:459; Gramentz 2008:140); 
“Cabinda” [-5.55000, 12.18333] (Frade 
1963:252), Turtle Taxonomy Working Group 
(2014:411). 
Taxonomic and distributional notes: 
Bour et al. (1995) discussed the type specimen. 
Map 119. Distribution of Cycloderma aubryi in Angola. 
Genus Trionyx Geoffroy Saint-Hilaire, 1809 
Trionyx triunguis (Forskal, 1775) Aerican Soetshell Turtle 
Testudo triunguis ForskM 1775: ix. Holotype, lost fide Webb in Iverson (1992), (collector P. Forskal). Type 
locality: “In Nilo” [= the Nile], Egypt. 
Gymnopus aegyptiacus: Bocage (1867b:218). 
