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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Trionyx triunguis: Peters (1877a:611), Boeage (1895a:7), Loveridge and Wiliams (1957:423), Loveridge 
(1957:172), Saldanha (1966:8), Hughes (1979:203), Iverson (1986:196, 1992:318), Braneh (1998:43,2008 
74), Turtle Taxonomy Working Group (2014:420). 
Amyda triunguis: Mertens (1926:152). 
Amyda triunguis triunguis: Hellmieh (1957a:33). 
Global conservation status (lUCN): Vulnerable. 
Global distribution: This species is widespread in tropical West Africa south of the Sahara as 
well as through the Nile Valley and around the eastern Mediterranean as far as Turkey. A disjunct 
population occurs in the Jubba and Scabelle rivers (Somalia), draining the Ethiopian Highlands. 
Ocurrences in Angola (Map 120): This 
species occurs in western Angola, mainly in 
near coastal drainages from Cabinda to the 
Cunene mouth. Cabinda: “Chinchoxo” 
[-5.10000, 12.10000] (Peters 1877a:611; 
Boeage 1895a:7; Loveridge and Wiliams 
1957:435). Bengo: “Cunga” [-9.23333, 
13.76667] (Loveridge and Wiliams 1957:435). 
Luanda: “Loanda”[-8.83333, 13.26667] 
(Loveridge and Wiliams 1957:435). Kwanza 
Norte: “Cuanza, Mucoso” [-9.53333, 
14.65000] (Hellmieh 1957a:33; Loveridge and 
Wiliams 1957:435). Benguela: “Lobito bay” 
[-12.35000, 13.55000] (Loveridge and Wil¬ 
iams 1957:435); “Catumbella” [-12.43333, 
13.55000] (Boeage 1867b:218, 1895a:7; 
Mertens 1926:152); Loveridge and Wiliams 
1957:435); “Benguella” [-12.58333,13.41667] 
(Boeage 1895a:7; Loveridge and Wiliams 1957:435). Namibe: 
11.80000] (Hughes 1979:203 [Fig. 67]). 
Taxonomic and distributional notes: Loveridge and Wiliams (1957) provided a detailed 
review of the species. 
Map 120. Distribution of Trionyx triunguis in Angola. 
Cunene mouth” [-17.28333, 
Order CROCODYLIA Gmelin, 1789 
Family Crocodylidae Cuvier, 1808 
Genus Crocodylus Laurenti, 1768 
Crocodylus niloticus Laurenti, 1768 Nile Crocodile 
Crocodylus niloticus Laurenti 1768:53. Type: not loeated fide King and Burke (1989:11). Type loeality: “Indi- 
en und Agypten”, later restrieted to “Agypten” [= Egypt] by Fuehs et al. (1974a: 110). 
Crocodilus vulgaris: Gunther (1865a:480), Boeage (1866a:41, 1867b:218, 1895a:8), Peters (1877a:611), 
Monard (1937b: 150). 
Crocodilus niloticus: Ferreira (1903:16), Themido (1941:11), Hellmieh (1957a:31), Saldanha (1966:8), 
Branch and McCarteney (1992:3), Ceriaco et al. (2014b:669), Branch and Conradie (2015:200), Conradie 
et al. (2016:26). 
Crocodylus niloticus chamses: Fuchs et al. (1974a:111). 
Global conservation status (lUCN): Least Concern. 
Global distribution: Widespread across much of eastern sub-Saharan Africa, northward along 
