MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
I8I 
I957a:36); “Chivitidi” [-13.01667, 14.63333] (Hellmich I957a:36); “Cubal” [-13.03333, 
14.25000] (Mertens I937a:7, I938a:43I; Loveridge 1947:404). Namibe: “Mossamedes” 
[-15.20000, 12.15000] (Bocage I867c:227, I895a:I5; Loveridge 1947:404; Laurent I964a:37; 
Ceriaco et al. 20I6a:54); “Pico Azevedo” [-15.55000, 12.51667] (Schmidt 1933:5-6; Barbour and 
Loveridge 1946:164; Loveridge 1947:398, 404; Marx 1959:466; Ceriaco et al. 20I6a:54); “Rio 
Coroca” [-15.78333,12.06667] (Bocage I887b:202; Loveridge 1947:404; Ceriaco etal. 20I6a:54); 
“Iona National Park, 9.65 km (by air) west-south-west of Espinheira” [-16.81200, 12.27126] 
(Ceriaco et al. 2016a:24); “Espinheira” [-16.78639, 12.35799] (Ceriaco et al. 2016a:24); “Omauha 
Lodge” [-16.19861, 12.40087] (Ceriaco et al. 2016a:24); “Iona National Park, Rio Curoca in Pedi- 
va Hot Springs area” [-16.28359, 12.56106] (Ceriaco et al. 2016b:24); “Namibe-Lubango road, 
road marker 59, 1.8 km west (by road) of Caraculo, on the north side of the road” [-15.01592, 
12.64239] (Ceriaco et al. 2016a:24); “Pico Azevedo” [-15.53400, 12.49197] (Ceriaco et al. 
2016a:24). 
Taxonomic and distributional notes: The genus Chondrodactylus was previously considered 
monotypic (C. angulifer) but Bauer and Lamb (2005) demonstrated that a clade of large-bodied 
geckos, the Pachydactylus bibronii group (Lamb and Bauer 2002), is sister to C. angulifer and 
moved these taxa into Chondrodactylus. Pachydactylus bibronii pulitzerae was originally 
described by Schmidt (1933) from “Pico Azevedo” in southern Angola and was long considered a 
synonym of R bibronii (Smith, 1846) [= Chondrodactylus bibronii] (Ceriaco et al. 2014a). Benyr 
(1995), in an unpublished thesis, clarified the distinction between the more temperate C. bibronii 
and a more tropical Chondrodactylus laevigatus (Fisher, Chondrodactylus turneri (Gray, 
1864) lineage and treated C. pulitzerae as a subspecies of Chondrodactylus laevigatus (see also 
Lamb and Basuer 2002). Recently, Heinz (2011) demonstrated that Chondrodactylus pulitzerae is 
a distinct, species-level lineage, occurring in sympatry or at least parapatry with C. laevigatus in 
southern Angola and in extreme northern Namibia. Most records for this species are in the south¬ 
west of Angola in Benguela Province (Schmidt 1933; Parker 1936; Monard 1937b; Mertens 1937a; 
Hellmich 1957a; Laurent 1964a), although Bocage (1895a) reported it to be common south of the 
Kwanza in general and cited two specimens from “Loanda” collected by Bayao in 1874. A recent 
record from Capanda Dam is a northward range extension of the species. It has also recently been 
recorded Cacuaco, Luanda Province (Ceriaco et al. 2017), suggesting that C. pulitzerae may be 
more widely distributed throughout the savanna biome of Angola (Ceriaco et al. 2014b). Although 
cited in the chresonymy, specific localities from the unpublished theses of Benyr (1995) and Heinz 
(2011) have not been listed above. 
Chondrodactylus laevigatus (Fischer, 1888) Button-Scaled Gecko 
Pachydactylus laevigatus (Fischer 1888:15, pi. 2, fig. 3). Syntypes: ZMH (not located) (collector J. Stein- 
grover). Type locality: “bei Aus und auf dem Wege nach Bethanien,” [= Aus on the way to Bethanien, 
Karas Region] Namibia. 
Pachydactylus stellatus: Schmidt (1933:5). 
Pachydactylus bibronii-. Monard (1937b:53). 
Pachydactylus bibronii turneri (part): Loveridge (1947:405). 
Paehydaetylus bibronii pulitzerae'. Hellmich (1957b:49) 
Paehydaetylus laevigatus laevigatus'. Benyr (1995:7). 
Paehydaetylus turneri (part): Branch (1998:254). 
Chondrodaetylus turneri (part): Heinz (2011:30), Bates et al. (2014:104), Ceriaco et al. (2014b:670). 
Chondrodaetylus cf pulitzerae (part): Conradie et al. (2016:24). 
Global conservation status (lUCN): Not Evaluated. 
