182 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Global distribution: The species is known from the north of the Northern Cape Province, 
South Africa, throughout Namibia and into southern Angola and north and east as far as southern 
Kenya. 
Ocurrences in Angola (Map 128): The 
species appears to occur in Cunene, Huila, and 
Cuando Cubango provinces, with a single 
questionable record fom southern Namibe 
Province. Huila: “Kampulu (pres Kasinga)” 
[-15.21667, 16.11667] (Monard 1937b:53; 
Loveridge 1947:407); “Mulondo” [-15.63333, 
15.20000] (Schmidt 1933:5; Loverige 
1947:407-409); “Gambos” [-15.76667, 
14.10000] (Hellmich 1957b:49). Cunene: 
“Humbi” [-16.68333, 14.90000] (Monard 
1937b:53; Loveridge 1947:407); “Mupanda” 
[-17.13333, 15.76667] (Monard 1937b:53; 
Loveridge 1947:407). Cuando Cubango: 
“Cuito basin (30a)” [-17.50875, 20.06594] 
(Conradie et al. 2016:9); “Cuito basin (62)” [- 
17.50875, 20.06608] (Conradie et al. 2016:9). 
Taxonomic and distribntional notes: , 128 . Distribution of cWmdacofc in 
Angola. 
The genus Chondrodactylus was previously 
considered a monotypic (C. angulifer) but Bauer and Lamb (2005) demonstrated that a clade of 
large-bodied geckos, the Pachydactylus bibronii group (Lamb and Bauer 2002), is sister to 
C. angulifer and moved these taxa into Chondrodactylus. Benyr (1995) clarified the distinction 
between the more temperate Chondrodactylus bibronii (Smith, 1846) and a more northerly distrib¬ 
uted Chondrodactylus 1. laevigatus (Fisher 1888) and Chondrodactylus 1. turneri (Gray, 1864). 
Branch (1998) subsequently used the name turneri as a senior synonym of laevigatus, correcting 
Benyr’s oversight of the former name’s priority (Lamb and Bauer 2002; Ceriaco et al. 2014b) when 
the two are treated as conspecific. Heinz (2011) demonstrated that C. turneri and C. laevigatus are 
deeply divergent from one another, but Heinicke et al. (2017) were the first to employ C. laeviga¬ 
tus as specifically valid subsequent to its transfer to Chondrodactylus. Schmidt (1933) reported 
P. stellatus [syn. Chondrodactylus laevigatus^ from Pico Azevedo (the type locality of P. pulitzer- 
ae), but these specimens are referable to C.fitzsimonsi (see account above). Loveridge (1947) erro¬ 
neously considered Monard’s (1937b) material of Pachydactylus bibronii from “Humbi,” “Kam- 
pulo,” and “Mupanda” representative of Chondrodactylus turneri, a species now not believed to 
occur in Angola. Although cited in the chresonymy, specific localities from the unpublished theses 
of Benyr (1995) and Heinz (2011) have not been listed above. 
Genus Hemidactylus Oken, 1817 
Hemidactylus angulatus complex Hallowell, 1852 African Tropical Gecko 
Hemidactylus angulatus Hallowell 1852a:63, fig. Holotype: ANSP 7431 (donated by H. Ford). Type loeality: 
“West eoast of Afriea” [= Gabon] by implieation fide Loveridge (1947). 
Hemidactylus brookii angulatus: Laurent (1964a:29). 
Hemidactylus brooki: Spawls et al. (2004:86). 
Hemidactylus angulatus: Carranza and Arnold (2006:539), Bauer et al. (2006a:8), Trape et al. (2012:222), 
Trape et al. (2012:222). 
