MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Pachydactylus amoenoides: Laurent (1964a:36). 
Pachydactyluspunctatus: Bauer and Braneh (1995a:70), Braneh (1998:259), Bauer et al. (2006b:646), Bauer 
(1999:57), Bates et al. (2014:130), Cerlaeo et al. (2016a:26, 55). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The speeies as presently eonstrued is widespread across southern Africa 
from the northwest and northeast of South Africa north to Malawi, the former Katanga Province of 
the Democratic Republic of Congo, and southern Angola. 
Ocurrences in Angola (Map 142): Pub¬ 
lished species records are from southwestern 
Angola, but members of this clade also occur in 
parts of central and southeastern Angola. 
Benguela: “Hanha” [-12.25000, 13.75000] 
(Bauer and Branch 1995a;82), “Lobito” 
[-12.33333, 13.50000] (Laurent 1954a:63, 
1964a:36; Bauer and Branch 1995a;82); 
“Catumbella” [-12.43333, 13.55000] (Bocage 
1895a: 16; Loveridge 1947:356); “Benguella” 
[-12.58333, 13.41667] (Bocage 18676:220, 
1895a; 16; Boulenger 1885:205; Loveridge 
1947:356). Huila: “Indungu” [-14.81667, 
16.26667] (Monard 1937b;54); “Capelongo” 
[-14.91667, 15.08333] (Bauer and Branch 
1995a;82); “Humpata, environs de Sa da Ban- 
deira” [-15.03333, 13.40000] (Laurent 
1964a;36); “Kului” [-15.41667, 15.73333] 
(Monard 193 7b: 5 4). Namibe: “ km 60 sur la 
route de Mogamedes a Sa da Bandeira” [-15.00000, 12.66667] (Laurent 1964a;36; Ceriaco et al. 
2016a:55); “11 miNE ofMocamede” [-15.08783, 12.26833] [Bauer and Branch 1995:82; Ceriaco 
et al. 2016a;55); “environs de Mogamedes” [-15.16667, 12.15000] (Laurent 1964a;36; Ceriaco et 
al. 2016a:55); “35 km south of Mogamedes” [-15.50000, 12.16667] (Laurent 1964a:36; Ceriaco et 
al. 2016a:55); “Pico Azevedo” [-15.55000, 12.51667] (Schmidt 1933:5; Loveridge 1947:356; 
Ceriaco et al. 2016a:55); “Iona National Park” [-16.65669, 12.43672] (Ceriaco et al. 2016a:26); 
“Espinheira” [-16.78639, 12.35799] (Ceriaco et al. 2016a;26). Cunene: “Riv. Mbale” [-15.16667, 
16.75000] (Monard 1931:90); “Forte Rogadas” [-16.71667, 15.01667] (Laurent 1964a:36). Unde¬ 
termined Locality: “Cuanza” (Boulenger 1905:110; Loveridge 1947:356); “Arid subregion” 
(Frade 1963:253). 
Taxonomic and distributional notes: Members of the Pachydactylus punctatus complex 
have been confused with a number of other southern African Pachydactylus, including the species 
P. geitje (Sparrman, 1778) and P. serval Werner, 1910, South African and Namibian endemics, 
respectively (Loveridge 1947; Bauer and Branch 1995a). Bocage’s (1867b, 1895a), Boulenger’s 
(1885, 1905) and Trade’s (1963) Angolan specimens of P. ocellatus (= P. geitje) and Monard’s 
(1931, 1937b) Pachydactylus serval specimens from “Mbale,” “Indungu,” and “Kului” have all 
been redetermined as P. punctatus (Loveridge 1947; Bauer et al. 2006b). Bauer and Lamb (2005) 
and Heinicke et al. (2011, 2017) identified P. punctatus as part of the “northwestern clade” of 
Pachydactylus, which comprises numerous morphologically diverse species that are widely dis¬ 
tributed in southern Angola and northern Namibia. Laurent (1964a) considered Pachydactylus 
amoenoides Hewitt, 1935 to be distinct from P. punctatus based on their sympatry in southern 
Map 142. Distribution of Pachydactylus punctatus in 
Angola. 
