MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Rhoptropus afer. Bocage (1897b:210). 
Rhoptropus ? boultoni: Parker (1936:127). 
Rhoptropus afer benguellensis: Mertens (1938a: 430, 431). 
Rhoptropus boultoni benguellensis: Mertens (1967:59), Hellmieh (1957a:37), Laurent (1964a:33), Bauer and 
Good (1996:643), Heinieke et al. (2017:6). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: Endemic to western Angola. 
Ocurrences in Angola (Map 149): The 
species is endemic to the southwest quadrant 
of Angola, north of the Namib. Benguela: 
“Bocoio” [-12.46667, 14.13333] (Parker 
1936:127); “Cubal” [-13.03333, 14.25000] 
(Mertens 1938a:431; Laurent 1964a:33; Mer¬ 
tens 1967:59; Bauer and Good 1996:643); 
“Entre Rios” [-13.01667, 14.63333] (Hellmieh 
1957b:37; Laurent 1964a:33); “Marco de 
Canavezes (Cubal de Ganda)” [-13.08333, 
14.33333] (Laurent 1964a:33); “Hanha” 
[-13.30000, 14.20000] (Bocage 1897b:210). 
Taxonomic and distributional notes: 
Three subspecies of Rhoptropus boultoni, 
including R. b. benguellensis, were long recog¬ 
nized (Bauer and Good 1996; Bauer and Lamb 
2001), but recent molecular phylogenetic 
11 1 j .1 .L 11 .Li -jy Map 149. Distribution of Rhoptropus benguellensis in 
research has revealed that all three are specifi- 
cally distinct (Kuhn 2016; Heinieke et al. 2017) ^ 
and do not form a monophyletic group. Loveridge (1947) and Laurent (1964) regarded a number 
of earlier references to other Rhoptropus to be applicable to this taxon, and we here have followed 
them. Although formally published localities are from Benguela, recent collections have included 
material from as far north as the Cuanza River in central Angola (Kuhn 2016). This species occu¬ 
pies some of the most mesic environments of any member of the genus. In the original description 
of Mertens (1938), the two images of the type specimen were mistakenly labeld as Rhoptropus afer 
benguellensis. 
Rhoptropus biporosus FitzSimons, 1957 FitzSimons’ Namib Day Gecko 
Rhoptropus biporosus FitzSimons 1957:395. Holotype: TM 24198 (collector Bernard Carp/Transvaal 
Museum Expedition). Type locality: “Ompemhe, 120 miles west of Ohopoho [= Opuwo], Kaokoveld, 
Southwest Africa,” Kunene Region, Namibia. 
Rhoptropus biporosus: Bauer and Good (1996:643), Branch (1998:267), Ceriaco et al. (2016a:27, 55). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: Recorded only from inland of the northern Namib dune fields form 
extreme northern Namibia and into adjacent Angola. 
Ocurrences in Angola (Map 150): The species is present in southern Namibe Province below 
the Escarpment. Namibe: “Pico Azevedo” [-15.53400, 12.49197] (Bauer and Good 1996:643; 
Ceriaco et al. 2016a:55); “Iona National Park” [-16.53347, 12.44560] (Ceriaco et al. 2016a:27); 
“Iona National Park, 20 km south-south-west (by air) of Espinheira” [-16.93169, 12.24500] 
