212 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Zygaspis nigra is sympatric with Z quadri- 
frons (Peters, 1862) in portions of its range 
(Broadley and Broadley 1997), although 
allopatrie in eastern Angola and Namibia. The 
typical habitats for this species are miombo 
woodlands and Baikiaea woodlands on sandy s 
Map 164. Distribution of Zygaspis nigra in 
Angola. 
Is (Broadley and Broadley 1997; Branch 1998). 
Zygaspis quadrifrons (Peters, 1862) Kalahari Round-Snouted Worm Lizard 
Global conservation status (lUCN): Not 
Evaluated. 
Ocurrences in Angola (Map 164): Pub¬ 
lished records in Angola derived from the 
northeastern provinces. Lunda Sul: “Alto Chi- 
capa” [-10.88333, 19.23333] (Laurent 
1964a: 84; Broadley and Gans 1969:1; 
Broadley and Broadley 1997:14). Moxico: 
“Gago Coutinho District” [-14.101654, 
21.435308] (Broadley and Gans 1969:1; 
Broadley and Broadley 1997:14; Broadley and 
Measey 2016:118, 122); “Colanda 7 km E of 
Vila Luso” (Broadley and Gans 1975:21; 
Broadley and Broadley 1997:14; Broadley and 
Measey 2016:118). 
Taxonomic and distributional notes: 
I3°E 16“E 20'=E 24°E 
Amphisbaena quadrifrons Peters 1862a:25. Syntypes: ZMB 4202-4204 (eolleetor C. H. Hahn). Type loeali- 
ty: “Neu-Barmen, im Hereolande, an der Westkiiste von Afrika” [= Gross Barmen, Otjozondjupa Region], 
Namibia. 
Amphisbaena ambuellensis Monard 1931:93, figs. 1-4. Syntype: MHNC 91.0621 (eolleetor A. Monard). Type 
loeality: “Chimporo” and “Caquindo” [= Tehimpolo and Caquindo], Cunene and Cuando Cubango 
provinees, Angola respeetively. 
Amphisbaena ambuellensis: Monard (1937b:65). 
Zygaspis quadrifrons quadrifrons: Loveridge 
(1941e:385), Gans (1967:88). 
Zygaspis quadrifrons: Broadley and Broadley 
(1997:14), Gans etal. (1997:44), Gans (2005:39), 
Conradie et al. (2016:23). 
Global distribution: A widespread 
species distributed from Southern Democratic 
Republic of Congo, south through Zambia, 
Zimbabwe, and Botswana to the northern 
regions of South Africa, west to southwestern 
Angola and northern Namibia, east to southern 
Malawi and central Mozambique 
Ocurrences in Angola (Map 165): The 
species has been reported from southern Ango¬ 
la, but may be expected to occur in Moxico 
Province near the border with Zambia as well. 
Cunene: “Chimporo” [-16.03333, 17.15000] 
Map 165. Distribution of Zygaspis quadrifrons in 
Angola. 
