MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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(Monard 1931:93, I937b:65; Loveridge I94Ic:385, 387; Gans 1967:88, 2005:39; Broadley and 
Broadley 1997:17). Cuando Cubango: “Caquindo (Kakindo)” [-15.45000, 17.05000] (Monard 
1931:93, I937b:65; Loveridge I94Ic:385, 387; Gans 1967:88, 2005:39; Broadley and Broadley 
1997:17); “lower Cuando river (41b)” [-17.46777, 23.07944] (Conradie et al. 2016:9, 11, 23); 
“lower Cuando river (44b)” [-17.56916, 23.27305] (Conradie et al. 2016:9, 11, 23); “Cuito basin 
(59)” [-16.28392, 18.84744] (Conradie et al. 2016:9, 12, 23); “Cuito basin (60)” [-16.92367, 
19.29675] (Conradie et al. 2016:9, 12, 23). 
Taxonomic and distributional notes: Bauer et al. (1995a) and Gans (2005) mistakenly gave 
the current name of Neu Barmen as Otjimbingue. This is, in fact, the location of another Rhenish 
mission station operated in the 19* century by the missionary and collector Carl Hugo Hahn. 
Amphisbaena ambuellensis (Monard, 1931) was synonymized with Z. quadrifrons with some mis¬ 
givings by Loveridge (1941c) because he expected to find “racial differentiation” between the 
fauna of southern Angola and that of South West Africa [Namibia] and Broadley and Broadley 
(1997) likewise considered that these populations might eventually prove to be distinct. 
Family Lacertidae Bonaparte, 1831 
Genus Heliobolus Fitzinger, 1843 
Heliobolus lugubris (Smith, 1838) Bushveld Lizard 
Lacerta lugubris Smith 1838:93. Syntypes: BMNH 1946.8.6.37-45 (collector A. Smith). Type locality: “Dis¬ 
trict immediately beyond the northern frontier of the [Cape] Colony,” South Africa. 
Eremias lugubris: Bocage (1867b:221, 1895a:31), Boulenger (1905:111; 1921:239), Monard (1937b:75), 
Mertens (1938a:437). 
Lampreremias lugubris: Szczerbak (1975:33). 
Heliobolus lugubris: Branch (1998:161), Bates et al. (2014:160), Ceriaco et al. (2016a:37, 56), Conradie et al. 
(2016:24). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: Widely distributed, from southern Angola over the Kalahari sands 
through Namibia, Botswana, southwestern Zimbabwe, southern Mozambique and South Africa. 
Ocurrences in Angola (Map 166): The 
species occurs broadly in southern Angola. 
Benguela: “Sighting in Lobito” [-12.35000, 
13.55000] (Monard 1937b:75); “Catumbella” 
[-12.43333, 13.55000] (Bocage 1895a:31); 
“Quissange” [-12.43333, 14.05000] (Bocage 
1895a:31); “Benguella” [-12.58333, 13.41667] 
(Bocage 1867b:221, 1895a:31; Szczerbak 
1975:33); “Dombe” [-12.95000, 13.10000] 
(Bocage 1895a:31); “Cubal” [-13.03333, 
14.25000] (Mertens 1938a:437); “Huxe, 
Benguella” [-12.71667, 13.20000] (Boulenger 
1921:243). Namibe: “Maconjo” [-15.01667, 
13.20000] (Bocage 1895a:31; Ceriaco et al. 
2016b:56); “Namibe-Lubango road, 2 km E of 
Mangueiras, south side of the road” 
[-15.04361, 13.16000] (Ceriaco et al. 
2016a:3 7); “Capangombe” [-15.10000, 
I2"E 20°E 24“E 
Map 166. Distribution of Heliobolus lugubris in 
Angola. 
