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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
13.15000] (Bocage 1895a:31; Ceriaco et al. 2016a:56); “Konondoto, Mossamedes” [-15.20000, 
12.15000] (Boulenger 1921:243; Ceriaeo et al. 2016a:56); “Namibe-Lubango road, 2 km E of 
Mangueiras, south side of the road” [-15.03333, 13.16000] (Ceriaco et al. 2016a:37); 
“Mossamedes” [-15.78333,12.06667] (Szczerbak 1975:33). Cunene: “PonangKuma” [-17.05000, 
14.65000] (Boulenger 1921:243); “Mupanda” [-17.13333, 15.76667] (Monard 1937b:75). Cuan- 
do Cubango: “Cuito basin (63)” [-17.93611, 21.10269] (Conradie et al. 2016:9, 12, 24). Unde¬ 
termined Locality: “Cuanza R.” (Boulenger 1905:111). 
Taxonomic and distributional notes: This widespread species has never been investigated in 
detail and it is likely that there is regional differentiation and possibly even cryptic taxa within 
H. lugubris as currently construed. 
Genus Holaspis Gray, 1863 
Holaspis guentheri Gray, 1863 
Blue-Tailed Tree Lizard 
Holaspis guentheri Gray 1863a:152, p. 20, fig. 1. Holotype: BMNH 1946.8.7.31 (donor Sir A. Smith, collec¬ 
tor unknown). Type locality: None stated, but Gray (1863) considered that it might be a “Tropical-Amer¬ 
ican” form. 
Holaspis guentheri: Ferreira (1897b:242), Spawls et al. (2004:163), Chirio and LeBreton (2007:218), Trape 
et al. (2012:322). 
Holaspis guentheri guentheri: Loveridge (1957:227), Laurent (1964a:56), Chirio and LeBreton (2007:218), 
Trape et al. (2012:322). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: This is a lowland forest species, found in open woodland adjacent forest 
and in some coastal forests, from Uganda and Tanzania in the east to Sierra Leone in the west. In 
the southwest it reaches Angola. 
Ocurrences in Angola (Map 167): The 
few published Angolan records are from the 
northeast of the country. Lunda Sul: “Alto 
Cuilo” [-10.01667, 19.55000] (Laurent 
1964a:56); “Alto Chicapa” [-10.88333, 
19.23333] (Laurent 1964a:56). 
Taxonomic and distributional notes: 
Based on the species global distribution record 
from “Rio Cuce,” Huila Province seems doubt¬ 
ful and is probably based on a misidentifica- 
tion. Ferreira’s (1897b) remarks relative to 
Gray’s (1863) and Boulenger’s (1887) descrip¬ 
tions of the species suggest that the “Rio Cuce” 
specimen was probably the small, terrestrial 
gerrhosaurid, Cordylosaurus subtessellatus 
(Smith, 1844), which shares a similar color pat¬ 
tern and does occur in the area. 
Map 167. Distribution of Holaspis guentheri in Angola. 
Genus Ichnotropis Peters, 1854 
Ichnotropis bivittata bivittata Bocage, 1866 Angolan Rough-Scaled Lizard 
lehnotropis Dumerilii [lehnotropis bivittatus] Bocage 1866a:43: Syntypes: ZMB 5827 fide Bauer et al. 
(1995b:41), MBL specimen numbers unknown, destroyed by fire 18 March 1978 (collector F.A.P. Bayao); 
Type loeality: “Duque de Bragan^a” (Boeage 1866a:43) [= Calandula], Malanje Provinee, Angola. 
