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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
“Rio Coroca” [-15.78333, 12.06667] (Bocage 1895a:32, 1897a:195; Boulenger 1887:111; Bauer 
and Gunther 1995:55; Ceriaco et al. 2016a:56); “Praia do Navio coastal dunes, ca. 124 km SSW of 
Namibe” [-16.27758, 11.82236] (Ceriaco et al. 2016a:36). 
Taxonomic and distributional notes: Bocage’s (1867a) doubts about the species’ inclusion 
in Scapteira were laid to rest by Strauch (1867). Boulenger (1887,1921) noted that one of the types 
of S. reticulata from “Benguella” was present in Berlin. Bauer and Gunther (1995) identified this 
specimen as ZMB 6475, and although not found at the time of their writing, it has since been locat¬ 
ed. However, although this specimen was collected by d’Anchieta and received from Bocage, it 
cannot be one of the types, as all five syntypes were from Mossamedes. Similarly to M. anchietae, 
there are very few published records for the species in Angola. 
Meroles squamulosus (Peters, 1854) Common Rough-Scaled Lizard 
Ichnotropis squamulosa Peters 1854:617. Syntypes: ZMB 6614, 6615, 6616 fide Bauer et al. (1995:58). Type 
loeality: “Tette” [= Tete], Mozambique. 
Ichnotropis squamulosa'. Monard (1937h:74), Loveridge (1957:234), Braneh (1998:163), Edwards et al. 
(2012:4), Engleder et al. (2013:132). 
Meroles squamulosus'. Edwards et al. (2013a: 101), Bates et al. (2014:164), Conradie et al. (2016:25). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: Meroles squamulosus occurs from Angola and Tanzania southwards 
through Zimbabwe, central Mozambique, Botswana, eastern Namibia, and northeastern South 
Africa. 
Ocurrences in Angola (Map 175): The 
species occurs in southern Angola east of the 
arid southwestern zone. Huila: “Kapelongo” 
[-14.88333, 15.08333] (Monard 1937b:74). 
Cunene: “Pereira de Eqa [= Ondijiva])” 
[-17.06666667, 15.73333333] (Conradie et al. 
2016:25). Cuando Cubango: “Cuito basin 
(30e)” [-17.51430, 20.05527] (Conradie et al. 
2016:9, 10,25); “Cuito basin (35)” [-16.62322, 
19.05352] (Conradie et al. 2016:9, 12, 25). 
Taxonomic and distributional notes: A 
recent analysis using nuclear and mitochondri¬ 
al sequence data revealed that this species, long 
known by the name Ichnotropis squamulosa, is 
strongly supported as a member of the genus 
Meroles (Edwards et al. 2012; Engleder et al. 
2013; Edwards et al. 2013a). Confusion with 
1 ^ . 1 7- 7 ^ • j Map 175. Distribution of Meroles squamulosus in 
members of the genus Ichnotropis is under- 
standable, as they are very similar morpholog¬ 
ically (Edwards et al. 2012; Edwards et al. 2013a). The geographic range and habitat of/, squa¬ 
mulosa overlaps with that of a number of Ichnotropis, but not that of other Meroles (Branch 1998). 
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