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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Ceriaco et al. 2016b:56); “ km 34 de lar oute de 
Mogamedes a Sa da Bandeira” [-15.50000, 
12.16667] (Laurent 1964a:56; Broadley 
1972:31; Ceriaeo et al. 2016b:5 6 ). Cunene: 
“Donguena, Mossamedes” [-17.01667, 
14.71667] (Boulenger 1910:472, 1917:210, 
1920:20; Broadley 1972:31). 
Taxonomic and distributional notes: 
Bocage (1895a) recorded Nucras from 
“Maconjo” and “Caconda” collected by Anchi¬ 
eta and identified them as belonging to N. tae- 
niolata (Smith, 1838). Boulenger (1910) also 
allocated his specimens from “Dongoena” 
(= Mossamedes fide Boulenger 1910) to this 
species, but subsequently (Boulenger 1917, 
1920) transferred them to N. holubi (Stein- 
dachner, 1882). Broadley (1972) revised the 
Nucras tessellata complex and recognised 
three subspecies: Nucras tessellata tessellata, 
Nucras tessellata livida (now a full species) and an unnamed subspecies of Nucras tessellata from 
Angola. Broadley also noted that the Angolan specimens were similar to N. taeniolata, but believed 
that more material was needed before the status of Angolan populations could be settled. Edwards 
et al. (2013b) confirmed the sister species relationship of N. tessellata to N. taeniolata, however, a 
detailed molecular investigation is still needed to resolve the position and status of Angolan 
Nucras, which may represent one or more new species (W.R. Branch, pers. comm., January 2017). 
Genus Pedioplanis Fitzinger, 1843 
Pedioplanis benguellensis (Bocage, 1867) Bocage’ s Sand Lizard 
Eremias benguellensis Bocage 1867h:221, 1867d:229. Holotype: MBL specimen number unknown (collector 
J.A. d’Anchieta), destroyed by fire 18 March 1978. Type locality: “Benguella” [= Benguela], Benguela 
Province, Angola. 
Eremias benguelensis: Boulenger (1921:287), Parker (1936:134). 
Eremias benguellensis: Monard (1937b:73), Laurent (1964a:60). 
Eremias sp.: Bocage (1887b:203). 
Eremias namaquensis: Boulenger (1887:91), Bocage (1895a:31). 
Eremias undata: Boulenger (1921:283). 
Eremias undata undata: Laurent (1964a:60). 
Mesalina benguelensis: Szczerbak (1975:24). 
Pedioplanis undata: Arnold (1989:213), Branch (1998:173), Makokha et al. (2007:623), Haacke (2008:90). 
Pedioplanis benguellensis: Arnold (1989:213), Bauer and Gunther (1995:55), Branch (1998:173), Makokha 
et al. (2007:623), Conradie et al. (2012b:93), Ceriaco et al. (2016a:37, 56). 
Pedioplanis namaquensis: Branch (1998:172), Makokha et al. (2007:623). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known from Angola and Nambia. 
Ocurrences in Angola (Map 178): The species occurs in the southwestern Angola. Benguela: 
“Catumbella” [-12.43333, 13.55000] (Bocage 1895a:31; Monard 1937b:73); “Benguella” 
[-12.58333, 13.41667] (Bocage 1867b:221, 1867d:229, 1895a:31; Monard 1937b:73; Szczerbak 
I2"E 1&“E 20“E 
Map 177. Distribution of Nucras aff. tessellata in 
Angola. 
