MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
223 
Map 178. Distribution of Pedioplanis benguellensis in 
Angola. 
1975:24; Bauer and Gunther 1995:55); “Huxe, 
Benguella” [-12.71667, 13.20000] (Boulenger 
1921:286, 288-289); “Catengue” [-13.03333, 
13.73333] (Parker 1936:134). Hulla: “Cacon- 
da” [-13.73333, 15.06667] (Bocage I895a:3I; 
Monard I937b:73). Namibe: “Namibe-Luban- 
go road, road marker 59, 1.8 km W (by road) 
from Caraculo, north side of the road” 
[-15.01606, 13.64483] (Ceriaeo et al. 
20I6a:37); “Maeonjo (=Fazendo Mueungo)” 
[-14.782192, 12.486557] (Boulenger 1921: 
286; Bauer and Gunther 1995:55; Conradie et 
al. 2012b:93; Ceriaeo et al. 2016a:56); “Capan- 
gombe” [-15.10000, 13.15000] (Boeage 
1895a:31; Monard 1937b:73; Ceriaeo et al. 
2016a:56); “Mossamedes” [-15.20000, 
12.15000] (Boeage 1887b:203, 1895a:31; 
Monard 1937b:73; Szezerbak 1975:24; Ceria- 
co et al. 2016a:56). Cunene: “Ponang Kuma” 
[-17.05000, 14.65000] (Boulenger 1921:289). 
Taxonomic and distributional notes: Boeage provided descriptions of this species in two dif¬ 
ferent papers, which appeared back-to-back in the same issue of the “Jomal de Sciencias Mathe- 
maticas, Physicas e Naturaes.” In Bocage (1867d) the description is short and benguellensis is 
spelled with two “l”s, whereas in Bocage (1867b) it is more extensive and the specific epithet is 
spelled with only one “1.” Although most authors have cited the more extensive description in 
Bocage (1867b), all except Boulenger (1921), Parker (1936) and Szezerbak (1975), including 
Bocage himself in later works, have adopted the spelling of the shorter description (Bocage 1867d). 
We here follow page priority and employ the prevailing spelling, benguellensis. Eremias benguel¬ 
lensis was later considered by Bocage (1895a) as a synonym of E. namaquensis Dumeril and 
Bibron, 1839. Boulenger (1921) reinstated Eremias. benguelensis as a full species, diagnosing it 
from E. namaquensis. This interpretation was subsequently followed by several authors, including 
Parker (1936), Monard (1937b) and Laurent (1964a), although 7^ benguellensis has frequently been 
synonymized with P. namaquensis. Based on phylogenetic analysis carried out by Conradie et al. 
(2012b), material referable to P. benguellensis does not fall within the same lineage as (nor is it sis¬ 
ter to) P. namaquensis, despite past speculation to the contrary (Bocage 1895a; Branch 1998; 
Makokha et al. 2007). Previous records of P. namaquensis from southern Angola should be con¬ 
sidered misidentifications, since there is no support for the occurrence of P. namaquensis in Ango¬ 
la (Conradie et al. 2012b). 
Pedioplanis haackei Conradie, Measey, Branch and Tolley, 2012 Haacke’s Sand Lizard 
(Endemic) 
Pedioplanis haackei Conradie et al. 2012b: 101, figs. 3A, B. Holotype: PEM R18465 (eolleetors W.R. Braneh, 
W. Conradie, G.J. Measey and K.A. Tolley) Type loeality: “along the road to Tambor, Namibe Provinee,” 
Namibe Province, Angola. 
Eremias undata undata (part): Laurent (1964a:60). 
Pedioplanis haackei'. Ceriaeo et al. (2016b:37, 56). 
Global conservation status (lUCN): Not Evaluated. 
