MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
227 
some time and until recently, C. miopropus was treated as a northern subspecies of C. macrolepis 
(Cope, 1862) (Broadley 1971d; Branch 1998; Spawls et al. 2004). Although this species is distin¬ 
guished by C. macrolepis by the presence of forelimbs (Bocage 1895a, Schmidt 1933; Loverdige 
1944b; Broadley 1971a; Broadley and Cotterill 2004) which are absent in the latter taxon. Chamae- 
saura miopropis is geographically isolated, occurring from Angola east to northern Malawi and 
southern Tanzania, whereas C. macrolepis is endemic to South Africa, Zimbabwe and Swaziland 
(Broadley and Cotterill 2004; Bates et al. 2014). Chamaesaura miopropus should therefore be con¬ 
sidered a valid species (Broadley and Cotterill 2004; Stanley et al. 2011; Bates et al. 2014). 
Genus Cordylus Laurenti, 1768 
Cordylus angolensis (Bocage, 1895) Angolan Girdled Lizard (Endemic) 
Zonurus angolensis Bocage 1895a:25. Holotype: MBL specimen number u nk nown (collector J.A. d’Anchi¬ 
eta), destroyed by fire 18 March 1978. Type locality: “Caconda,” Huila Province, Angola. 
Zonurus cordylus: Boulenger (1897:277). 
Zonurus angolensis: Monard (1937b:61), Frade (1963:252). 
Cordylus cordylus angolensis: Loveridge (1944b:38). 
Cordylus tropidosternum tropidosternum: Broadley (1971:22) 
Cordylus angolensis: Broadley and Branch (2002:10), Branch et al. (2005:131), Stanley et al. (2011:67), 
Greenbaum et al. (2012:36), Nielsen and Colston (2014:167), Stanley et al. (2016:23). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known from Angola. 
Ocurrences in Angola (Map 183): The 
species is known from the type locality 
“Caconda” in Huila Province but its distribu¬ 
tion range reaches the southern regions of 
Kwanza Sul Province. Kwanza Sul: “Mombo- 
lo” [-11.91667, 14.85000] (Greenbaum et al. 
2012:36). Huila: “Caconda” [-13.73333, 
15.06667] (Bocage 1895a:24; Monard 
1937b:61; Loveridge 1944b:39). 
Taxonomic and distributional notes: In 
the description of Cordylus angolensis Bocage 
(1895a) expressed some doubts about the true 
identity of C. angolensis, leading him to name 
it in a “provisional manner.” The species was 
not cited in any further Bocage works after the 
description. With the destruction of the type 
specimen in the Lisbon museum fire, the taxo¬ 
nomic identity of the species became even 
more problematic. Broadley (1971b) consid¬ 
ered it a synonym of Cordylus tropidosternum (Cope, 1869). Loveridge (1944b) and Branch et al. 
(2005) noted that Bocage’s type description of C. angolensis lacked detail, although it is now con¬ 
sidered a valid species endemic to central Angola (Broadley and Branch 2002; Stanley et al. 2011; 
Nielsen and Colston 2014; Stanley et al. 2016). Greenbaum et al. (2012) found three Cordylus 
specimens in the collections of the American Museum of Natural History assignable to 
C. angolensis from “Mombolo”, approximately 190 km north-northwest of the type locality. These 
specimens agree with the original holotype description in some measurments and coloration. 
Map 
Angola. 
183. Distribution of Cordylus angolensis 
