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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
although they differ in the scale counts of longitudinal rows of ventral scales (Greenbaum et al. 
2012). Angolan Cordylus are currently being revised and the systematic position of C angolensis 
will hopefully be resolved. Cordylus angolensis was described within the account of Cordylus 
cordylus (Bocage 1895a). The two specimens treated as C. cordylus were from the Sociedade de 
Geografia de Lisboa, Portugal, and had only the locality “Angola.” Monard (1937b) considered 
these specimens to be attributable to Cordylus vittifer (Reichenow, 1887), although Loveridge 
(1944b) considered this opinion untenable and considered them assignable to C. angolensis. The 
identity of these specimens, which have recently been re-discovered remains uncertain, but is under 
study. 
Cordylus machadoi Laurent, 1964 Machado’s Girdled Lizard 
Cordylus vittifer machadoi Laurent 1964a:49, fig. 12. Holotype: MD 1840-1 (collector A. Barros Machado). 
Type locality: “Leba, Humpata, environs de Sa da Bandeira, Alt. 1800 m, Huila” [= Serra da Leba, Luban- 
go], Huila Province, Angola. 
Cordylus vittifer machadoi'. Broadley (1971b:22) 
Cordylus machadoi'. Branch (1998:195), Stanley et al. (2011:67), Greenbaum et al. (2012:23), Bates et al. 
(2014:192), Nielsen and Colston (2014:171), Stanley et al. (2016:202). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known from southwestern Angola and northwestern 
Namibia. 
Ocurrences in Angola (Map 184): The 
species is known from the type locality in 
Huila Province, however the species distribu¬ 
tion may extend to Namibe Province. Huila: 
“Leba, Humpata, environs de Sa da Bandeira” 
[-15.06667, 13.60000] (Laurent 1964a:49); 
“Humpata” [-14.96167, 13.34972]; 
[-15.17750, 13.32139E (Stanley et al. 2011:57; 
Stanley et al. 2016:209); “Nascente do Tchi- 
viuguira” [-15.16667, 13.30000] (Stanley et al. 
2016:209). 
Taxonomic and distributional notes: 
The holotype of this taxon has recently been 
rediscovered in the collection at Dundo. The 
single juvenile paratype, was exchanged from 
this collection and is now MCZ R-74120 (fide 
Stanley et al. 2016). Branch (1998) elevated 
C. machadoi to full species, an action recently 
Map 184. Distribution of Cordylus machadoi in Angola. 
supported by recently Stanley et al. (2016) who based their conclusions on two C. machadoi spec¬ 
imens collected at the type locality. These were recovered as sister to all East African species of 
Cordylus (Stanley et al. 2011; Greenbaum et al. 2012; Nielsen and Colston 2014). A cryptic species 
allied to C. machadoi, was recently discovered at lower elevation (Stanley et al. 2016). Significant 
genetic variation across such a constrained geographic area suggests that its northern congener, 
C. angolensis (Bocage, 1895), which has a comparatively broad geographic distribution, may itself 
represent a species complex (Stanley et al. 2016). 
