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Cordylus namakuiyus Stanley, Ceriaco, Bandeira, Valerio, Bates and Branch, 2016 
Kaokoveld Girdled Lizard (Endemic) 
Cordylus namakuiyus (Stanley et al. 2016:212, figs. 1, 4, 5). Holotype: CAS 254912 (collectors L. Stanley, 
S. de Sa, S. Bandeira, H. Valerio, A. Kuhn, J. Vindum and L. Ceriaco). Type locality: “large rock outcrop 
near Caraculo, road from Lubango and Namibe, Namibe Province,” Angola. 
Cordylus namakuiyus: Ceriaco et al. (2016a:38, 59). 
Stanley et al. (2016) recently described this 
form as a new species endemic to the arid low¬ 
lands west of the southern Angolan escarpment 
and sister to the geographically proximate, 
high-elevation species Cordylus machadoi 
Laurent, 1964. All phylogenetic analyses recovered the Angolan specimens as monophyletic, form¬ 
ing the earliest diverging lineage of the northern Cordylus clade (Stanley et al. 2016). These authors 
analyzed the morphology of AMNH Angolan specimens identified as Cordylus cordylus (Linnaeus, 
1758) collected during the Vemay expedition in 1925 and showed that they cluster closely with 
C. namakuiyus. Although the Vemay specimens lack specific locality information, the expedition 
field notes mention that significant numbers of unidentified lizards were collected at “Pico Azeve- 
do” and “100 km east of Mogamedes,” the same areas where the eight specimens of the 
C. namakuiyus were collected, leading the authors to assign the Vemay material to the new species 
(Stanley et al. 2016). 
Map 185. Distribution of Cordylus namakuiyus in 
Angola. 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known only from Angola. 
Ocurrences in Angola (Map 185): The 
species is known from Huila and Namibe 
provinces in southwestern Angola. Huila: 
“large rock outcrop near Caraculo, on the road 
from Lubango and Namibe” [-15.01650, 
12.64203] (Stanley et al. 2016:212; Ceriaco et 
al. 2016a:38, 59). Namibe: “outskirts of Pico 
Azevedo” [-15.47589,12.46269] (Stanley et al. 
2016:212; Ceriaco et al. 2016a:38, 59); “low 
rock outcrop bordering the road between 
Namibe and Omahua lodge” [-15.99681, 
12.40683] (Stanley et al. 2016:212; Ceriaco et 
al. 2016a:38, 59). 
Taxonomic and distributional notes: 
20“E 24''E 
Family Gerrhosauridae Fitzinger, 1843 
Genus Cordylosaurus Gray, 1865 “1866” 
Cordylosaurus subtessellatus (Smith 1844) Dwarf Plated Lizard 
Gerrhosaurus subtessellatus Smith 1844: pi. 41, fig. 2 and first of two accompanying unnumbered text pages, 
pi. 42, figs. 17-20. Holotype: BMNH 1946.8.19.41 (eolleetor A. Smith). Type loeality: “Great Namaqua- 
land” [= southern Namibia]. 
Cordylosaurus trivittatus: Boulenger (1887:126), Boeage (1867b:222, 1895a:37). 
Cordylosaurus subtessellatus: Ceriaeo et al. (2016a:39, 59). 
