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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
2007). According to Bates et al. (2013) G. bulsi is a distinct species that is sister to the clade {Ger- 
rhosaurus intermedius [G. auritus, G. nigrolineatus]). 
Gerrhosaurus multilineatus Bocage, 1866 Kwanza Keeled Plated Lizard (Endemic) 
Gerrhosaurus multilineatus Bocage 1866a:44. Holotype: MBL specimen number unknown (collector F.A.P. 
Bayao), destroyed by fire 18 March 1978. Type locality: “Duque de Braganga” [= Calandula], Malanje 
Province, Angola. 
Gerrhosaurus multilineatus-. Bocage (1866b:61, 1867b:221), Peters (1881:147). 
Gerrhosaurus bulsi'. Ceriaco et al. (2016b:63). 
Bocage (1866b) remarked that Gerrhosaurus 
multilineatus might be a well-characterized 
variety of G. nigrolineatus Hallowell, 1857, 
however he considered the two species distinct 
based on color pattern. Loveridge (1942) and 
FitzSimons (1943) subsequently relegated 
G. multilineatus to the synonymy of G. nigro¬ 
lineatus, the former considering it a color variant (Loveridge 1942). Haagner et al. (2000) stated 
that Broadley (unpublished) considered G. multilineatus to be based on a hybrid specimen and that 
the name would, therefore, be unavailable. However, according to Article 17.2 of the Code (ICZN 
1999), even if the specimen was a hybrid, the name would still be available (Bates et al. 2013). 
Currently this species is recognized as valid despite the obvious taxonomic confusion, the loss of 
the original type series to fire at the Museu Bocage and the lack of museum material, especially 
from Angola (Bates et al. 2013). The status of Gerrhosaurus multilineatus and the assignment of 
Angolan populations referred to Gerrhosaurus nigrolineatus (e.g., Boulenger 1887, 1905; Ferreira 
1900a, 1903, 1904; Angel 1923; Monard 1937b; Mertens 1938a; Hellmich 1957a, 1957b; Manaqas 
1963; Parker 1936; Schmidt 1933; Laurent 1954a) remains problematic and resolution must await 
the collection of material from the type locality for molecular analysis, and a detailed morpholog¬ 
ical evaluation of the complex (Bates et al. 2013). Ceriaco et al. (2016b) referred to the specimen 
collected in Cangandala National Park (Malanje Province) as G. bulsi, but ongoing molecular 
works reveal that it shall be considered true multilineatus. 
Map 189. Distribution of Gerrhosaurus multilineatus in 
Angola. 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known only from Angola. 
Ocurrences in Angola (Map 189): Pub¬ 
lished records exist for this species from cen¬ 
tral and western Angola. Malanje: “Duque de 
Braganqa” [-9.10000, 15.95000] (Bocage 
1866a;44, 1866b:61); “Malanje” [-9.55000, 
16.35000] (Peters 1881:147); “Cangandala 
National Park” [-9.84606, 16.72233] (Ceriaco 
et al. 2016b:63). Benguela: “Catumbella” 
[-12.43333, 13.55000] (Bocage 1867b;221); 
“Dombe” [-12.95000, 13.10000] (Bocage 
1867b:221). 
Taxonomic and distributional notes: 
20“E 
24''E 
