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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
[-11.18333, 17.45000] (Schmidt 1933:11); “Chitau” [-11.43333, 17.15000] (Schmidt 1933:11). 
Huambo: “Bimbi” [-11.81667, 15.83333] (Monard 1937b:78); “Galanga” [-12.06667, 15.15000] 
(Bocage 1895a:35; “Santo Amaro” [-12.70000, 15.85000] (Monard 1937b:78); “Cuma” 
[-12.86667,15.06667] (Loverdige 1936a:65). Benguela: “Cahata” [-12.35000, 14.81667] (Bocage 
1887c:210); “Catumbella” [-12.43333, 13.55000] (Bocage 1887c:210); “Quissange” [-12.43333, 
14.05000] (Bocage 1887c:210, 1895a:35); “Quindumbo” [-12.46667, 14.93333] (Bocage 
1895a:35); “Entre Rios” [-13.01667, 14.63333] (Hellmich 1957a:54); “Cubal” [-13.03333, 
14.25000] (Mertens 1938:435); “Benguela” [-12.58333, 13.41667] (Boulenger 1887:122; Bocage 
1895a:35); “Hanha” [-13.30000, 14.20000] (Bocage 1896a:lll); “Catengue” [-13.03333, 
13.73333] (Mertens 1926:152). Hulla: “Caconda” [-13.73333, 15.06667] (Bocage 1895a:35); 
“Kuvangu” [-14.46667, 16.30000] (Monard 1937b:78); “Kapelongo” [-14.88333, 15.08333] 
(Monard 1937b:78); “Quillengues” [-14.06667, 15.08333] (Bocage 1895a:35); “Huilla” 
[-15.05000,13.55000] (Bocage 1895a:35); “Kampulu” [-15.21667,16.11667] (Monard 1937b:78); 
“Mulondo” [-15.63333, 15.20000] (Monard 1937b:78). Namibe: “Capangombe” [-15.10000, 
13.15000] (Bocage 1895a:35; Ceriaco et al. 2016a:59). Cunene: “riv. Mbale” [-15.16667, 
16.75000] (Monard 1937b:78); “Chimporo” [-16.03333, 17.15000] (Monard 1931:109, 
1937b: 148). Cuando Cubango: “Kakindo” [-15.45000, 17.05000] (Monard 1937b:78); “Kwito 
region, tributary of Kubango” (Angel 1923:159); “Cubango basin (6a)” [-14.67155, 17.73525] 
(Conradie et al. 2016:8-9, 24); “Cuito basin (30a)” [-17.50875, 20.06594] (Conradie et al. 2016:9- 
10,24); “Cuito basin (35)” [-16.62322, 19.05352] (Conradie et al. 2016:9-10,24); “Cubango basin 
(46)” [-14.58981, 16.907389] (Conradie et al. 2016:9, 12, 24). Undetermined locality: “Carangi- 
go” (Bocage 1895a:35); “Between, Bihe and Quilenges” (Boulenger 1905:111); “without precise 
location” (Bocage 1870:68; Ferreira 1900:49). 
Taxonomic and distributional notes: Bates et al. (2013) suggested that G. nigrolineatus was 
not monophyletic and resurrected G. intermedius Lonnberg, 1907 for eastern populations (Kenya, 
Uganda, Rwanda, Tanzania, Malawi, Mozambique, Zimbabwe and South Africa). However, accu¬ 
rate determination of geographical boundaries for nigrolineatus and intermedius, especially in Cen¬ 
tral Africa (Angola, Democratic Republic of Congo, Zambia, northern Botswana and northern 
Namibia) will require additional sampling, as well as additional morphological analysis of speci¬ 
mens from throughout their extensive ranges. Bates et al. (2013) questioned the assignment of 
Angolan Gerrhosaurus multilineatus Bocage, 1866 specimens to G. nigrolineatus (e.g., Boulenger 
1887, 1905; Ferreira 1900, 1903, 1904; Angel 1923; Monard 1937b; Mertens 1938; Hellmich 
1957a, 1957b; Managas 1963; Parker 1936; Schmidt 1933; Laurent 1954), and their relationship 
(see G. multilineatus account). The status of the G. nigrolineatus complex in Angola is currently 
under investigation (M. Bates, pers. comm.) and the records plotted here should be considered as 
only tentatively assigned to this species. Gerrhosaurus nigrolineatus ahlefeldti (Hellmich and 
Schmelcher, 1956) is now considered a synonym of G. nigrolineatus, the holotype and four 
paratypes are present in the Zoological Museum in Hamburg and two paratypes are at the Zoolo- 
gische Staatssammlung, Miinchen (Hallermann 1998; Franzen and Glaw 2007). 
Gerrhosaurus skoogi Andersson, 1916 Desert Plated Lizard 
Gerrhosaurus skoogi Andersson 1916:10, fig. 2. Holotype: GNM 1387 (donor Lt. D. Fortunato). Type loeal- 
ity: “Port Alexander, Portuguese West Afriea” [= Tombua], Namibe Provinee, Angola. 
Angolosaurus skoogi'. Loveridge (1942:495), FitzSimons (1953:215), Braneh (1998:177), Ceriaeo et al. 
(2016a:39, 59). 
Global conservation status (lUCN): Least Concern. 
Global distribution: A deserticolous species that may form loosely-structured colonies 
