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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Taxonomic and distributional notes: 
The species was for some time considered a 
subspecies of Gerrhosaurus validus (Smith, 
1849). A combination of molecular, morpho¬ 
logical and geographical evidence, suggests 
that the two taxa represent separate evolution¬ 
ary lineages (Bates et al. 2013) that occur in 
allopatry, with validus distributed in southeast¬ 
ern Africa and maltzahni restricted to northern 
Namibia and southern Angola (Bates et al. 
2013; Bates et al. 2014). Thus records of Ger¬ 
rhosaurus validus (or Gerrhosaurus robustus 
Peters, 1854) from Angola (Bocage 1870, 
1887b, 1895a; Boulenger 1905; Mertens 
1938a; Frade 1963) are, in fact, referrable to 
maltzahni. A new genus, Matobosaurus was 
erected by Bates and Tolley (2013) to incorpo¬ 
rate this distinctive clade of gerrhosaurs (Bates 
et al. 2013). 
ll^E 16°E 20°E 24'*E 
Map 192. Distribution of Matobosaurus maltzahni in 
Angola. 
Genus Tetradactylus Merrem, 1820 
Tetradactylus ellenbergeri (Angel, 1922) Ellenberger’s Plated Snake-Lizard 
Paratetradactylus Ellenbergeri (Angel 1922:151). Holotype: MNHN 1921-514 (donated by V. Ellenberger). 
Type loeality: “Pays des Barotses (Rhodesie)” [= Barotseland] Zambia. 
Tetradaetylus lundensis (Monard 1937b:79). Syntypes: MHNC 91.0455-58 [4 speeimens] (eolleetor 
A. Monard). Type loeality: “Lunda, sur les bords du Tyihumbwe” [= Lunda, on the edges of Tyihumbue 
river] Lunda Sul Provinee, Angola. 
Caitia afrieana\ Boeage (1895a:37). 
Tetradaetylus lundensis (Monard 1937b:79). 
Tetradaetylus boulengeri lundensis: de Witte and i 2 »e i&°e 2 o°e 24 ”e 
Laurent (1942b: 105). 
Tetradaetylus ellenbergeri ellenbergi: Laurent 
(1964a:55). 
Tetradaetylus ellenbergeri boulengeri: Laurent 
(1964a:55). 
Tetradaetylus ellenbergeri: Broadley and Cotterill 
(2004:43), Wagner et al. (2012:35), Conradie et 
al. (2016:24). 
Global conservation status (lUCN): Not 
Evaluated. 
Global distribution: The species is 
known from Angola, Democratic Republic of 
Congo, Tanzania and Zambia. 
Ocurrences in Angola (Map 193): The 
species occurs in norteastem Angola. Lunda 
Norte: “Dundo” [-7.36667, 20.83333] (Lau- 
rent 1964a:55). Lunda Sul: “Lunda” Ua? m. m.Xnhutkyr, of TetraJactylus ellenbergeri in 
[-10.96667, 20.06667] (de Witte and Laurent Angola. 
