MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
281 
Loveridge 1947:208). Huambo: “Bela-Vista (= Sanguengue)” [-12.36667, 16.20000] (Hellmich 
1957b:53); “Mt. Moco” [-12.41667, 15.18333] (Parker 1936:141); “Cuma” [-12.86667, 15.06667] 
(Loveridge 1936a:76). Benguela: “Quibula” [-12.28333, 14.68333] (Bocage 1895a:59); “Cahata” 
[-12.35000, 14.81667] (Bocage 1895a:59); “Catumbella” [-12.43333, 13.55000] (Bocage 
1867b:219); “Quissange” [-12.43333, 14.05000] (Bocage 1887b:209, 1895:59); “Quindumbo” 
[-12.46667, 14.93333] (Bocage 1895a:59); “Benguella” [-12.58333, 13.41667] (Bocage 1866a:42, 
1866c:59,1867b:219,1895a:59-60; Loveridge 1957:200); “Dombe” [-12.95000,13.10000] (Boca¬ 
ge 1867b:219, 1895a:60); “Entre Rios/Chitidi” [-13.01667, 14.63333] (Mertens 1938a:435, Hell¬ 
mich 1957a:52); “Cubal” [-13.03333, 14.25000] (Mertens 1937a:7, 1938a:435; Hellmich 
1957b:53); “Marco de Canavezes, perto da barragem do rio Cubal” [-13.30000, 14.20000] (Boca¬ 
ge 1896a:112); “Hanha” [-13.30000, 14.20000] (Bocage 1896a:112); “Ebanga” [-12.73333, 
14.73333] (Monard 1937b:99). Hulla: “Caconda” [-13.73333, 15.06667] (Bocage 1879c:88; 
Loveridge 1936a:76); “Lubango” [-14.91667, 13.50000] (Bocage 1895a:59); “Huilla” [-15.05000, 
13.55000] (Bocage 1895a:59); “Mutundo” [-15.23333, 13.36667] (Frade 1963:231); “Gambos” 
[-15.76667, 14.10000] (Bocage 1895a:59); “Kampulu, environs de Kasinga” [-15.21667, 
16.11667] (Monard 1937b:99); “Kuvangu” [-14.46667, 16.30000] (Monard 1937b:99); “Kului” 
[-15.41667, 15.73333] (Monard 1937b:99). Namibe: “Mossamedes” [-15.20000, 12.15000] 
(Bocage 1867b:219, 1887b:202, 1895a:59; Boulenger 1887:451; Ceriaco et al. 2016a:58); “Chim- 
ba” [-15.20000, 13.68333] (Hellmich 1957b:53; Ceriaco et al. 2016a:58). Cunene: “Mupanda” 
[-17.13333, 15.76667] (Monard 1937b:99). Cuando Cubango: “Lwankundu, sub-tributary of 
Kwando” [Undetermined Locality] (Angel 1923:165); “Cuito basin (30a)” [-17.50875, 20.06594] 
(Conradie et al. 2016:9-10, 24). Undetermined locality: “Between Bihe and Quilenges” 
(Boulenger 1905:112); “Mayumba/Maiumba” (Bocage 1887a:178, 1895a:60, 1897a:198); “Cuan- 
za r.” (Boulenger 1905:112). Without precise locality: “with no precise locality” (Laurent 
1964a:44). 
Taxonomic and distributional notes: Bocage (1866a) noted some distinct characters in a 
specimen collected by Anchieta in “Rio Quilo” relative to the typical form, Chamaeleo dilepis 
Leach, 1819, however, variation across the C. dilepis species complex as a whole suggests that 
there are few if any consistent differences among these forms and the taxonomy of the C. dilepis 
complex remains unresolved (Glaw 2015). The issue is exacerbated by the vague type locality of 
C. dilepis, “Congo,”which, in the BMNH register is given as “Gaboon, French Congo” (although 
Pauwels and Vande weghe 2008) stated that the species is absent from Gabon). Loveridge (1936a) 
refered all Angolan specimens from the Field Museum to C. d. quilensis, with the exception of one 
specimen from “Cuma,” which he regarded as typical dilepis (FMNH 15365). Broadley (1966) and 
Tilbury (2010) considered quilensis a variant of the polymorphic C. dilepis, although Klaver and 
Bohme (1986) and Ullenbruch et al. (2007) identified differences in hemipenial morphology and 
regarded quilensis as subspecifically (Klaver and Bohme 1987) or even specifically distinct from 
nominotypical C. dilepis (Klaver and Bohme 1986). Tilbury (2010) provided an extended discus¬ 
sion of the taxonomic and distributional issues surrounding the C. dilepis complex. Regardless if 
quilensis is taxonomically distinct from C. dilepis sensu stricto, it seems likely that all members of 
the complex form Angola are referable to the same form (although some authors have assigned 
some Angolan material to another doubtful form, C. d. roperi) and we here treat all Angolan spec¬ 
imens as C. d. quilensis pending a more detailed phylogeographic study of the group. 
