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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Chamaeleo gracilis etiennei Schmidt, 1919 Etienne’s Graceful ChamaeleoN 
Chamaeleo gracilis Hallowell 1842:324, pi. 18. Holotype: ANSP 7325 (“purchased by the Rev. Charles Eden 
of Monrovia, of one of the Affiean natives”). Type loeality: “Liberia, Western Affiea.” 
Chamaeleon etiennei Sehmidt 1919:574. Holotype: AMNH 11370 (eolleetor H. Lang). Type loeality: “hills 
near Banana at the mouth of the Congo,” Demoeratie Republie of Congo. 
Chamaeleon graeilis: Boeage (1866a:41, 1867b:219, 1895a:61), Ferreira (1900a:50, 1903:16, 1904:117), 
Boulenger (1887:448, 1905:111), Monard (1937b:98), Themido (1941:8) 
Chamaeleon senegalensis: Boeage (1870:68), Peters (1881:147). 
Chamaeleon graeilis: Boulenger (1887:448, 1905:111), Boeage (1895a:61). 
Chamaeleo senegalensis: Gunther (1865a:480). 
Chamaeleon etiennei: Parker (1936:140), Laurent (1964a:42). 
Chamaeleo graeilis: Hellmieh (1957a:53, 1957b:53), Laurent (1964a:44). 
Chamaeleo graeilis graeilis: Loveridge (1957:198). 
Chamaeleo graeilis etiennei: Loveridge (1957:197). 
Chamaeleo senegalensis senegalensis: Loveridge (1957:197). 
Chamaeleon graeilis etiennei: Tilbury (2010:507), Ceriaeo et al. (2014b:670), Glaw (2015:202) 
Global conservation status (lUCN): Least Concern. 
Global distribution: Known from the sounthem Congo to northern Angola 
Ocurrences in Angola (Map 242): Limit¬ 
ed to areas from Kwanza Sul and Lunda Sul 
northwards, including Cabinda. Although we 
located no published historical records from 
Lunda Norte, Tilbury (2010) plotted them 
there, and it is likely that they occur in this 
province as well as the immediately adjacent 
Democratic Republic of Congo. Cabinda: 
“Chinchoxo” [-5.10000, 12,10000] (Peters 
1877a:612; 1881:147; Boeage 1895a:61); 
“Cabinda” [-5.55000, 12.18333] (Boeage 
1895a:61). Lunda Sul: “Alto Cuilo” 
[-10.01667, 19.55000] (Laurent 1964a:42); 
“Alto Chicapa” [-10.88333, 19.23333] (Lau¬ 
rent 1964a;42). Luanda: “Loanda” [-8.83333, 
13.26667] (Boeage 1867b:219). Kwanza 
Norte: “Piri-Dembos” [-8.56667, 14.50000] 
(Hellmieh 1957b:53); “Ambaca” [-9.26667, 
15.18333] (Boeage 1895a;61); “N’Dalla 
Tando” [-9.30000, 14.91667] (Ferreira 1904:111); “Cazengo” [-9.33333, 14.76667] (Ferreira 
1903:16); “Mucoso” [-9.53333, 14.65000] (Hellmieh 1957a;53); “Dondo” [-9.68333, 14.43333] 
(Boeage 1895a:61); “Canhoca” [-9.25000, 14.68333] (Ferreira 1904:111; Parker 1936:140). 
Lunda Sul: “Lunda” [-10.96667, 20.06667] (Monard 1937b:98); “Mutianvo” [-11.45000, 
19.33333] (Themido 1941:8). Mexico: “environs du lac Calundo” [-11.80000, 20.86667] (Laurent 
1964a:42). Kwanza Sul: “Lembu, Serra de Selles” [-12.86667, 14.11667] (Ferreira 1904:117); 
“Congulu” [-10.86667, 14.28333] (Parker 1936:140); “Condo” [-11.28333, 14.33333] (Gunther 
1865a:480; Boulenger 1887:448; Parker 1936:140). Benguela: “Ebanga” [-12.73333, 14.73333] 
(Monard 1937b:98). Undetermined locality: “Quango = Cuango” (Peters 1877a:612, 1881:147; 
Boeage 1895a:61) (Malanje Province, impossible to georeferenced. See History Section for more 
detailed information); “Carangigo” (Boulenger 1887:448). 
Map 242. Distribution of Chamaeleo gracilis etiennei in 
Angola. 
