MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
287 
Genus Agama Daudin, 1802 
Agama aculeata Merrem, 1820 Western Ground Agama 
Agama aculeata Merrem 1820:53. Leetotype: Speeimen illustrated in Fig. 6, plate 8 of Seba (1735) (eollee- 
tor unknown), probably lost, designated by Wagner et al. (2012). Type loeality: “Promontorio bonae spei” 
[= the Cape of Good Hope] South Afriea. 
Agama aculeata'. Boeage (1866a:43, 1867b:221), Braneh (1998:212), Leaehe et al. (2009:274), Wagner et al. 
(2012:177), Ceriaeo et al. (2016a:58). 
Agama armata: Boeage (1870:68, 1879e:88, 1895a:19, 1896a:110, 1896b:127), Boulenger (1905:110). 
Agama hispida aculeata'. Loveridge (1936a:52), Mertens (1938a:432), Hellmieh (1957a:38, 1957b:51), 
Mananas (1963:229), Laurent (1964a:40). 
Agama hispida var. aculeata'. Monard (1937b:59). 
Agama aculeata aculeata'. Bates et al. (2014:303). 
Global conservation status (lUCN): Least Concern. 
Global distribution: Endemic to southern Africa where it has an extensive range from south¬ 
ern Angola, most of Namibia except the Namib Desert proper, western Botswana and the western 
half of South Africa. 
Ocurrences in Angola (Map 247): The 
species widespreasd in Angola except for the 
more northerly provinces. Malanje: “Duque 
de Braganga” [-9.10000, 15.95000] (Boeage 
1866a:43, 1895a:19, 1896b:127). Lunda Sul: 
“Alto Chicapa” [-10.88333, 19.23333] (Lau¬ 
rent 1964a:40). Moxico: “Lago Cameia” 
[-11.71667, 20.80000] (Mananas 1963:229), 
“environs du lac Claundo” [-11.80000, 
20.86667] (Laurent 1964a;40), “Calombe, 
Luso” [-11.83333, 19.93333] (Mana 9 as 
1963:229); “Cassamba” [-13.10000, 20.35000] 
(Managas 1963:229). Bie: “Silva Porto” 
[-12.38333, 16.95000] (Mananas 1963:229); 
“Cubango basin (15)” [-12.78555, 16.75694] 
(Conradie et al. 2016:8-9, 23). Huambo: 
“Bimbi” [-11.81667, 15.83333] (Monard 
1937b:59); “Galanga” [-12.06667, 15.15000] 
(Boeage 1895a:19); “Bela-Vista (Sanguengue)” [-12.36667, 16.20000] (Hellmieh 1957b:51); 
“Nova Lisboa” [-12.76667, 15.73333] (Mananas 1963:229); “Santo-Amaro” [-12.70000, 
15.85000] (Monard 1937b:59); “Cuma” [-12.86667, 15.06667] (Loveridge 1936a:52). Benguela: 
“Cahata” [-12.35000, 14.81667] (Boeage 1895a: 19) “Quissange” [-12.43333, 14.05000] (Boeage 
1895a:19); “Benguella” [-12.58333, 13.41667] (Loveridge 1936a:52); “Ebanga” 
[-12.73333, 14.73333] (Monard 1937b:59); “Entre Rios” [-13.01667, 14.63333] (Hellmieh 
1957a:38); “Cubal” [-13.03333, 14.25000] (Mertens 1938:432); “Hanha” [-13.30000, 14.20000] 
(Boeage 1896:110). Huila: “Rio Cuce” [-13.51667, 15.20000] (Boeage 1895a:19); “Cassoco” 
[-13.65000, 15.01667] (Boeage 1895a:19); “Caconda” [-13.73333, 15.06667] (Boeage 1879c:88); 
“Kalukembe” [-13.78333, 14.68333] (Monard 1937b:59); “Kuvangu” [-14.46667, 16.30000] 
(Monard 1937b:59); “Boca de Humpata, Sa da Bandeira” [-14.93333, 13.51667] (Laurent 
1964a:40); “Chibia, Rio Huila” [-15.20000, 13.68333] (Hellmieh 1957b:51); “Kambisa” 
[-15.31667, 16.21667] (Monard 1937b:59); “Kului” [-15.41667, 15.73333] (Monard 1937b:59). 
Map 247. Distribution of Agama aculeata in Angola. 
