MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
309 
Genus Bids Gray, 1842 
Bids arietans (Merrem, 1820) Puff Adder 
Vipera {Echidna) arietans Merrem 1820:152. Leetotype: speeimen deseribed and illustrated by Seba 
(1735:55, pi. 54, fig. 4), now lost (eollector unknown), designated by Wallaeh et al. (2014:611). Type loeal- 
ity: “Vorgeburge der guten Hoffnung/Promontorio bonae spei” (Merrem 1820:152), [= Cape of Good 
Hope] Western Cape Provinee, Southern South Afriea. 
Echidna arietans: Boeage (1866a:53). 
Clotho arietans: Gunther (1865a:480). 
Echida arietans: Boeage (1879e:89). 
Bids arietans: Boeage (1887a:190, 1887e:211), Ferreira (1897b:245), Boulenger (1915:221), Sehmidt 
(1933:15), Loveridge (1936a:45), Monard (1937b: 141-142), Themido (1941:11), Hellmieh (1957a:75), 
Spawls and Braneh (1995:114), Braneh (1998:114), Lenk et al. (1999:31), Dobiey and Vogel (2007:104), 
Barlow et al. (2013:1134), Wallaeh et al. (2014:89), Braneh and Conradie (2015:200), Ceriaeo et al. 
(2016a:87), Conradie et al. (2016:23). 
Vipera arietans: Boeage (1895a: 149, 1896a: 113). 
Cobra lachesis: Mertens (1937a: 16). 
Bids lachesis: Mertens (1938a:442), Bogert (1940:99), Laurent (1950a: 11, 1954a:62). 
Bids arietans arietans: Hellmieh (1957b:74), Laurent (1964a: 127), Thys van den Audenaerde (1966:36), 
Mananas (1981-82:35), Branch and McCartney (1992:2), Bates et al. (2014:331). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The speeies is known from most of sub-Saharan Afriea, extending to the 
Arabian Peninsula. 
Ocurrences in Angola (Map 269): The 
speeies oeeurs in the entire eountry with 
exeeption of the desert regions of the far south¬ 
west, as well as some mountainous areas. 
Zaire: “S. Salvador do Congo” [-6.26667, 
14.23333] (Boeage 1887a:190, 1895a;149; 
Managas 1981:35). Bengo: “Ambriz” 
[-7.844312, 13.106493] (Managas 1981:35). 
Kwanza Norte: “Mucoso, Dondo” [-9.53333, 
14.65000] (Hellmieh 1957a:75; Mananas 
1981:35). Malanje: “Duque de Bragan^a” 
[-9.10000, 15.95000] (Boeage 1866a:53, 
1895a: 149; Monad 1937b: 141; Managas 
1981:35); “Cangandala National Park” 
[-9.84606, 16.72233] (Ceriaeo et al. 
2016a:87). Lunda Norte: “Dundo” [-7.36667, 
20.83333] (Laurent 1950a:ll, 1954a:62, 
1964a: 127; Mananas 1981:35); “Capaia” 
[-7.33556, 20.21681] (Braneh and Conradie 2015:200); “Muita, Luembe E” [-7.80000, 21.45000] 
(Laurent 1950a: 11, 1954a:62; Managas 1981:35); “Sombo” [-8.68333, 20.95000] (Laurent 
1954:62); “Maludi” [-8.03333, 21.33333] (Thys van den Audenaerde 1966:36). Lunda Sul: “Alto 
Chieapa” [10.93333,19.15000] (Laurent 1964a:127; Managas 1981:35). Moxico: “environs du lae 
Calundo (Lago Calundo)” [-11.80000, 20.86667] (Laurent 1964a:127; Managas 1981:35). Bie: 
“Chitau” [-11.25000, 17.16667] (Sehmidt 1933:15; Bogert 1940:99; Managas 1981:35); “Cuban¬ 
go basin (9)” [-14.00269, 17.40500] (Conradie et al. 2016:8-9, 23). Huambo: “Bela Vista” 
[-12.56667, 16.21667] (Managas 1981:35; Hellmieh 1957b:74); “Huambo” [-12.83333, 15.66667] 
Map 269. Distribution of Bids arietans in Angola. 
