MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Global distribution: The species is known from western Angola, chiefly in association with 
the high plateau. 
Ocurrences in Angola (Map 272): The 
species is known from the central highlands in 
Angola. Zaire: “S. Salvador do Congo” 
[-6.26667, 14.23333] (Managas 1981:38). 
Bengo: “Bengu” [-8.71667, 13.40000] 
(Monard 1937b:141; Mertens 1958:147; 
Managas 1981:38; Spawls and Branch 
1995:105; Wallach et al. 2014:91). Kwanza 
Sul: “Mombolo” [-12.16667, 14.83333] 
(Bogert 1940:101; Mertens 1958:146; Mana- 
gas 1981:37-38). Huambo: “Bela Vista (San- 
guengue)” [-12.56667, 16.21667] (Hellmich 
1957b:75; Mertens 1958:146; Mana 9 as 
1981:38; Spawls and Branch 1995:105). Bie: 
“Bie” [-12.38333, 16.95000] (Mertens 
1958:147; Managas 1981:38; Spawls and 
Branch 1995:105) Huila: “Caconda” 
[-13.73333, 15.06667] (Bocage 1895a:151; 
Map 272. Distribution of Bitis heraldica in Angola. 
Ferreira 1897b:245; Monard 1937b:141; Mertens 1958:147; Managas 1981:38; Spawls and Branch 
1995:105); “Kalukembe (Caluquembe)” [-13.78333, 14.68333] (Monard 1937b:141, 143; Mertens 
1958:147; Mananas 1981:38; Spawls and Branch 1995:105); “sur les bords de la riviere Calae, I’un 
des affluents du Cunene (Rio Calae)” [-13.96667, 16.03333] (Bocage 1889:127, 1895a:151; 
Monard 1937b:141; Mertens 1958:147; Managas 1981:38; Spawls and Branch 1995:105). Unde¬ 
termined Locality: “Between Benguella and Bihe” (Boulenger 1905:114; Monard 1937b: 141). 
Taxonomic and distributional notes: This species was described by Bocage (1889) based on 
one specimen from “sur les bords de la riviere Calae, Tun des affluents du Cunene”. This species 
was later considered by Boulenger (1896) a synonym of Bitis peringueyi (Boulenger, 1888) 
although Ferreira (1897) doubted this conclusion. Later, Mertens (1958) restablished B. heraldica 
based on distinctive morphological characters between the two species and refered Angolan mate¬ 
rial from “Kalukembe” (Monard, 1937b) and from “Namba, Mombolo” (Bogert, 1940) to 
B. heraldica. 
Bitis nasicornis (Shaw, 1792) Rhinoceros Viper 
Coluber Nasicomis: Shaw in Shaw and Nodder 1792: pi. 94, first of three aeeompanying unnumbered pages 
of Latin deseription). Holotype: BMNH, lost fide Golay et al. (1993) (donated by E. Jenkins). Type loeal- 
ity: “interiori Afriea” [“likely to be from Ghana”fide Hughes and Barry 1969:1030]. 
Bitis nasieornis: Boulenger (1915:222), Parker (1936:126), Hellmieh (1957b:76), Loveridge (1957:302), 
Mananas (1981:36), Spawls and Braneh (1995:118), Lenk et al. (1999:31), Lenk et al. (2001:96), Chirio 
and LeBreton (2007:604), Dobiey and Vogel (2007:117), Wallaeh et al. (2014:91). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is restricted to the rainforest of West Africa, known from 
Guinea to Gabon, extending east to Uganda, western Kenya, and northern Tanzania, having its 
southern limit in the northern regions of Angola and Zambia. Wallach et al. (2014) considered 
records from Spawls et al. (2004) from northeastern Tanzania to be unconfirmed. 
Ocurrences in Angola (Map 273): The species is limited to the northwestern of the country. 
