MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
315 
“Tomboa,” is the first explicit published record for the species in the country (Spawls and Branch 
1995). Wallach et al. (2014) erroneously considered B. peringueyi present in Kwanza Sul Province, 
presumably based on Bogert’s (1940) record of 5. heraldica (as “5. peringueyi”). 
Genus Causus Wagler, 1830 
Causus bilineatus Boulenger, 1905 Two-Striped Night Adder 
Causus rhombeatus var. bilineatus Boulenger 1905:114. Syntypes: BMNH 1905.5.29.36-40 (eolleetor W.J. 
Ansorge). Type loeality: “Duque de Braganea, Quissange, Caeonda et Huilla” (Boeage 1895a: 146) 
[= Calandula, Quissange, Caeonda and Huila] Angola, and “Between Benguela and Bihe” (Boulenger 
1905:114), Angola. 
Causus rhombeatus-. Boeage (1895a:147); Boulenger (1896:468); Sehmidt (1933:15), Hellmieh (1957b:74), 
Loveridge (1957:299). 
Causus bilineatus bilineatus'. Laurent (1964a: 125). 
Causus bilineatus'. Spawls et al. (2004:469), Broadley (1968b:407), Spawls and Braneh (1995:140), Broadley 
and Cotterill (2004:46), Rasmussen (2005:1), Mananas (1981:33), Dobiey and Vogel (2007:125), Wallaeh 
et al. (2014:148). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known from the plateau of South-Central Africa, in Ango¬ 
la, southern Democratic Republic of Congo and north-west Zambia, reaching Tanzania. 
Ocurrences in Angola (Map 275): The 
species occurs from west to eastern Angola. 
Kwanza Norte: “Piri-Dembos” [-8.53333, 
14.43333] (Hellmieh 1957b;74; Manaqas 
1981:33; Rasmussen 2005:14); “Canhoca” 
[-9.25000, 14.68333] (Boulenger 1905:114; 
Manaqas 1981:33). Malanje: “Duque de Bra- 
ganqa” [-9.10000, 15.95000] (Boeage 
1866a:51, 1895a;145; Monard 1937b;141, 
Manaqas 1981:33; Rasmussen 2005:14). 
Moxico: “Rives du lac Calundo (Lago Calun- 
do)” [-11.80000, 20.86667] (Laurent 
1964a;125; Manaqas 1981:33; Rasmussen 
2005:14). Bie: “Chitau” [-11.43333,17.15000] 
(Schmidt 1933:15; Rasmussen 2005:14). 
Huila: “Caeonda” [-13.73333, 15.06667] 
(Boeage 1895a:145; Monard 1937b:141; 
Mana?as 1981:33; Rasmussen 2005:14); Map 275 Distribution of 6, in Angola. 
“Capelongo” [-14.88333, 15.08333] (Manapas 1981:33); “Huila” [-15.08333, 13.55000] (Boeage 
1895a:145; Boulenger 1896:468; Monard 1937b:141; Manaqas 1981:33; Rasmussen 2005:14). 
Taxonomic and distributional notes: Boeage (1895a) in his remarks on Causus rhombeatus 
(Lichtenstein, 1823) recognized a striped form based on several individuals from Angola deposit¬ 
ed in the Museu Boeage collection from “Duque de Braganqa, Quissange, Caeonda et Huilla,” 
although he did not recognize them as taxonomically distinct. Boulenger (1905) subsequently 
noted the same striped form described by Boeage based on specimens from “between Benguella 
and Bihe” and proposed the name C. rhombeatus var. bilineatus. McDiarmid et al. (1999) includ¬ 
ed Boeage’s (1895a) specimens among the type series, but Boulenger (1905) clearly refered only 
to the striped specimens collected by Ansorge and in the BMNH collection. For some time, the 
