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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
nomen bilineatus was neglected (e.g., Schmidt 1933; Monard, 1937b; Bogert, 1940; Hellmich 
1957a,b) and the corresponding Angolan records were associated with C rhombeatus. Laurent 
(1964a) revived the name bilineatus, and included his recently described Causus lineatus (Laurent, 
1956), which he considered a dwarf subspecies of Causus bilineatus. However, his specimens 
differ little from those of the type series from “Benguela to Bihe” (Broadley and Cotterill 2004). 
Loveridge (1957) included both bilineatus and lineatus in the synoymy of C rhombeatus, but 
Broadley (1968b) placed C b. lineatus in the synonymy of C bilineatus without comment, where 
it has since remained (Rasmussen 2005). Laurent (1964a) erroneously assigned some Angolan 
records of C. rhombeatus to C bilineatus (e.g., “Pungo-Andongo” [Boulenger 1905]; “Mombolo” 
[Bogert 1940]; “Bela Vista”, “Entre Rios” [Hellmich 1957b]) as did Managas (1981). Most of the 
specimens cited by Laurent (1964a) were examined and revised by Rasmussen (2005). 
Causus lichtensteinii (Jan, 1859) Forest Night Adder 
A[spidelaps] (Sepedon) Lichtensteinii Jan 1859:511. Holotype: NMBA, lost (fide Kramer 1978; Golay et al. 
1993), destroyed in 1943 during the World War II fide Wallaeh et al. (2014). Type loeality: “Cote-d’Or” 
[= Ghana]. 
Causus liehtensteinii: Laurent (1950a:ll, 1964a:127), Mananas (1981:34), Spawls and Braneh (1995:143), 
Spawls et al. (2004:471), Rasmussen (2005:11), Chirio and LeBreton (2007:606), Dobiey and Vogel 
(2007:126), Akani et al. (2012:192), Wallaeh et al. (2014:149). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known from the central African forests from Guinea and 
Liberia eastward through Ghana to Nigeria, Cameroon, the Central African Republic, south to 
Democratic Republic of Congo, northeastern 
Angola, and northwestern Zambia, and east to 
Uganda and western Kenya. 
Ocurrences in Angola (Map 276): The 
species occurs in the northtem Angola, in the 
adjacent areas near the border with Democratic 
Republic of Congo and in the Cabinda encalve, 
although there are no published records from 
Cabinda. Lunda Norte: “Dundo” [-7.36667, 
20.83333] (Laurent 1964a: 127, Mananas 
1981:34); “riviere Lukashi, afluente de la rive 
gauche de la Tshihumbwe (Rio Lukashi)” 
[-7.53333, 21.08333] (Laurent 1950a:ll; 
Mananas 1981:34). 
Taxonomic and distributional notes; 
Wallaeh et al. (2014) recorded this species as 
present in Benguela Province, although this 
almost certainly is based on a misidentifica- 
tion. 
Map 
Angola. 
276. Distribution of Causus lichtensteinii in 
Causus maculatus (Hallowell, 1842) Spotted Night Adder 
Distiehurus Maeulatus Hallowell 1842:337, pi. 19. Holotype: ANSP 6897 (eolleetor S.M.E. Goheen). Type 
loeality: “Liberia, Westen Afriea”. 
Causus rhombeatus-. Boeage (1895a:145), Parker (1936:126), Laurent (1950a:ll, 1954a:61), Hellmieh 
(1957b:74). 
Causus maeulatus-. Laurent (1964a:124), Thys van den Audenaerde (1966:36), Managas (1981:32), Spawls 
