MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
317 
and Branch (1995:145), Rasmussen (2005:14), Chippaux (2006:241), Trape and Mane (2006:206), Dobiey 
and Vogel (2007:127), Chirio and LeBreton (2007:608), Wallach et al. (2014:148). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known from West-Central Africa from Mauritania to 
Ethiopia, and south to Angola. 
Ocurrences in Angola (Map 277): The 
species occurs in the northern Angola. Lunda 
Norte: “Dundo” [-7.36667, 20.83333] (Lau¬ 
rent 1950a: 11, 1954a;61, 1964a: 124, Thys van 
den Audenaerde 1966:36, Managas 1891:32; 
Rasmussen 2005:14); “Barrage de la Luachi- 
mo” [-7.38333, 20.85000] (Thys van den 
Audenaerde 1966:36); “Muita (Luembe E)” 
[-7.80000, 21.45000] (Laurent 1950a:ll, 
1954a;61); “Sombo” [-8.68333, 20.95000] 
(Laurent 1954a:61). Kwanza Norte: “Piri- 
Dembos” [-8.53333, 14.43333] (Hellmich 
1957b:74; Rasmussen 2005:14). Kwanza Sul: 
“Quirimbo” [-10.68333, 14.26667] (Parker 
1936:126). 
Taxonomic and distributional notes; 
Bogert (1940) pointed out a possible taxonom¬ 
ic difference between the savanna C. rhombea- 
tus (typical form) and the forest dwellers, which was later confirmed by Laurent (1956, 1964a). 
Laurent (1960) identified C. maculatus as a forest form, distinguishing it from C. rhombeatus, 
which is sympatric in Angola (Mana 9 as, 1981). Laurent (1964a) assigned the Angolan records pre¬ 
viously refered to C. rhombeatus (e.g., “Dundo,” “Muita,” “Sombo” [Laurent 1950a, 1954a], 
“Quirimbo” [Parker 1936]) to C. maculatus, and refered the specimens cited in Bocage (1895a) 
from Congo and a part of northern Angolan to C. maculatus. Without further comment. According 
to Rasmussen (2005) two specimens from the Zoological Museum of the University of Hamburg 
previously identified as Causus rhombeatus (Lichtenstein, 1823) from “Piri-Dembos” (Hellmich, 
1957b) correspond, in fact, to Causus maculatus. 
Map 277. Distribution of Causus maculatus in Angola. 
Causus resimus (Peters, 1862) Green Night Adder 
Heterophis resimus Peters 1862b:277, pL, figs. 4, 4a, 4b. Holotype: ZMB 4370 (eolleetor A. von Bamin and 
R. Hartman). Type loeality: “Sennar, vom Gebel-Ghule” [= Jebel Ghule, Sennar Provinee], Sudan. 
Heterophis resimus = Causus rostratus: Bocage (1870:68). 
Causus resimus: Bocage (1887c:211), Ferreira (1904:116), Boulenger (1896:469, 1915:220), Loveridge 
(1936a:44, 1957:299), Parker (1936:127), Monard (1937b:141), Bogert (1940:97), Managas (1981:33), 
Spawls and Branch (1995:142), Spawls et al. (2004:474). Rasmussen (2005:14), Chirio and LeBreton 
(2007:610), Dobiey and Vogel (2007:128), Akani et al. (2012:189), Wallach et al. (2014:149). 
Causus resimus var. angolensis Bocage (1895a: 148). Syntypes: MBL 2025, 2080 (Biballa), MBL 2078, 
2084a-b (Quissange), MBL 2079 (Novo Redondo), MBL 2082a-b (Rio Chimba), 2083a-b (Dondo) (fide 
McDiarmid et al. 1999) and presumably others, (collectors Banyures [Rio Dande, Rio Bengo], F.A.P. 
Bayao [Dondo], A. da Fonseca [Cazengo], F. Newton [Novo Redondo], J.A. d’Anchieta [Quissange, Rio 
Chimba, Biballa, Maconjo]), destroyed by fire 18 March 1978. Type locality: “Rio Dande et Rio Bengo ... 
Dondo ... Cazengo ... Novo Redondo ... Quissange, Rio Chimba, Biballa et Maconjo.” 
Causus rhombeatus: Hellmich (1957b:74). 
