MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
339 
Map 299. Distribution of Lycophidion multimaculatum in 
Angola. 
1968:474); “Calonda” [-8.41667, 20.53333] 
(Laurent I964a:94; 1968:474); “Lunda Dis¬ 
trict” (Broadley 1996b: 19). Lunda Sul: “Alto 
Cuilo” [-10.01667, 19.55000] (Laurent 
1964a:95; 1968:474). Mexico: “Cazombo” 
[-11.88333, 22.91667] (Laurent 1964a:95; 
1968:474); “Macondo, Haut Zambeze” 
[-12.55000, 23.76667] (Laurent 1964a:95; 
1968:474). Kwanza Norte: “Piri-Dembos” 
[-8.56667, 14.50000] (Ferreira 1906:167; 
Broadley 1996b: 19); “Golungo Alto” 
[-9.13333, 14.76667] (Ferreira 1906:167). Bie: 
“32 km E of Dande, Gauca” [-11.18333, 
17.45000] (Schmidt 1933:13; Broadley 
1996b:19); “Chitau” [-11.43333, 17.15000] 
(Schmidt 1933:13; Laurent 1968:474; 
Broadley 1996b: 19). Huambo: “Bela-Vista” 
[-12.36667, 16.20000] (Hellmich 1957b:61; 
Broadley 1996b:19); “Galanga” [-13.73333, 
15.06667] (Bocage 1895a:81; Laurent 1968:474). Benguela: “Entre-Rios” [-12.26667, 17.41667] 
(Hellmich 1957a:61, 71; Broadley 1996b: 19); “Hanha” [-13.30000, 14.20000] (Bocage 
1896a:112). Hufla: “Caconda” [-13.73333, 15.06667] (Bocage 1895a:81; Boulenger 1893:616; 
Laurent 1968:474; Broadley 1996b:19); “Rio Cuce” [-13.51667, 15.20000] (Ferreira 1897b:243; 
Monard 1937b:113); “Kalukembe” [-13.78333, 14.68333] (Monard 1937b:117); “Gambos” 
[-15.76667, 14.10000] (Broadley 1996b:19); “28 km N of Sa da Bandeira” [-14.916667, 
13.499914] (Broadley 1996b: 19). Namibe: “Mossamedes” [-15.20000, 12.15000] (Bocage 
1895a:81; Laurent 1968:474; Broadley 1996b:19). Cuando Cubango: “approximately 50 km E of 
Cuito Cuanavale” [-15.23333, 19.61667] (Branch and McCartney 1992:1; Broadley 1996b: 19). 
Undetermined Locality: “Between Benguela and Bihe” (Boulenger 1905:112; Broadley 
1996b: 19). 
Taxonomic and distributional notes: The majority of the specimens originally assigned to 
Lycophidion capense ^Smith, 1831, in Angola (Peters 1877a, 1881, Bocage 1895a, 1896a; Ferreira 
1904, 1906; Boulenger 1893, 1905, Schmidt 1933; Monard 1937b:117; Bogert 1940; Hellmich 
1957a, 1957b) refer to Lycophidion multimaculatum Boettger, 1888. Modem authors (e.g., 
Broadley 1996b; Branch 1998) have typically followed Broadley (1992b) in recognizing L. multi¬ 
maculatum as a full species. Given that the currently understood species boundary between this 
species and L. capense is suspiciously consistent with political boundaries, further investigation of 
populations of wolf snakes in and adjacent to the Caprivi Strip in northeastern Namibia is required. 
Lycophidion ornatum Parker, 1936 Ornate Wolf Snake 
Lycophidion ornatum Parker 1936:122. Holotype: BMNH 1946.1.14.28 (formerly BMNH 1936.8.1.690) (eol- 
leetor K. Jordan). Type loeality: “Congulu” [= Congulo], Kwanza Sul Provinee, Angola. 
Lycophidion capense ornatum'. Loveridge (1957:252). 
Lyeophidion ornatum'. Laurent (1968:470), Broadley (1992b:35, 1996b: 16), Spawls et al. (2004:325), 
Broadley and Hughes (1993:12), Chippaux (2006:80), Chirio and LeBreton (2007:474), Wallaeh et al. 
(2014:411). 
Global conservation status (lUCN): Least Concern. 
