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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Genus Prosymna Gray, 1849 
Prosymna ambigua Bocage, 1873 East African Shovel-Snout 
Prosymna ambiguus Bocage 1873b:218. Holotype: MBL 610 (collector F.A.P. Bayao), destroyed by fire 18 
March 1978. Type locality: “Duque de Braganga” [= Calandula], Malanje Province, Angola. 
Prosymna ambigua brevis Laurent 1954a:50, figs. 12-14. Holotype: MD 2177 (collector A. Barros Macha¬ 
do), lost (see Notes below). Type locality: “Dundo,” Lunda Norte Province, Angola. 
Prosymna meleagris: Bocage (1866a:47). 
Prosymna ambigua: Boulenger (1893:248, 1915:208), Bocage (1895a:99), Loveridge (1933:244), Monard 
(1937b: 114), Wallach et al. (2014:568), Ceriaco et al. (2016b:83). 
Prosymna bocagii: Boulenger (1915:208). 
Prosymna ambigua brevis: Laurent (1964a:108), Thys van den Audenaerde (1933:34), Loveridge (1958:157). 
Prosymna ambigua ambigua: Laurent (1950a:9, 1954a:52), Loveridge (1958:151), Broadley (1980:534), 
Chippaux (2006:142). 
Global conservation status (lUCN): Least Concern. 
Global distribution; The species is known from central Cameroon, Gabon and southern 
Congo, east through the southern half of Democratic Republic of Congo, northern Angola and 
Zambia to South Sudan, southwestern Kenya and Uganda and south to Malawi and Zimbabwe. 
Ocurrences in Angola (Map 303): The 
species occurs in the northern regions of the 
country. Lunda Norte: “Dundo” [-7.36667, 
20.83333] (Laurent 1950:9, 1954a:50, 
1964a: 108; Loveridge 1958:158; Thys van den 
Audenaerde 1966:34; Broadley 1980:538); 
“Sombo” [-8.68333, 20.95000] (Laurent 
1954:50; Loveridge 1958:158; Broadley 
1980:538). Malanje: “Duque de Braganqa” 
[-9.10000, 15.95000] (Bocage 1866a:47, 
1873a:218, 1895a:99; Loveridge 1933:244, 
1958:151-154; Monard 1937b:114; Broadley 
1980:538; Chippaux 2006:142; Wallach et al. 
2014:568); “Cangandala National Park” 
[-9.84606,16.72233] (Ceriaco et al. 2016b:83). 
Taxonomic and distributional notes: 
Broadley (1980) synonymized the subspecies 
P. a. brevis Laurent, 1954 with the typical Map 303. Distribution of Prosymna amNgwa in Angola. 
form. In January 2017 the holotype of P. a. brevis could not be located in the Museu Dundo. 
Prosymna angolensis Boulenger, 1915 Angola Shovel-Snout 
Prosymna angolensis Boulenger 1915:208. Syntypes: MBL specimen numbers unknown, lost fide Broadley 
(1980). Neotype: NHMW 19275:2 designated by Broadley (1980:497). Type locality: Angola by implica¬ 
tion (Boulenger 1915:208), restricted to “Huila” by Loveridge (1958:149), changed to “Caconda” by 
subsequent designation of a neotype Broadley (1980:497, 513). 
Prosymna frontalis: Bocage (1873b:218, 1895a:98, pi. 11, fig. 2), Boulenger (1893:248, 1896:641). 
Prosymna ambigua: Monard (1931:104, 1937b: 123). 
Prosymna angolensis: Monard (1937b: 114, 122), Loveridge (1958:149), Bogert (1940:59), Hellmich 
(1957b:66), FitzSimons (1962:161), Broadley (1980:512, 1990:227), Branch (1998:84), Broadley et al. 
(2003:187), Wallach et al. (2014:568). 
Prosymna ambigua ambigua: Mertens (1938a:439), Loveridge (1958:151). 
