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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
“Dondo, Luanda” (Crawford-Cabral and Mesquitela 1989). Psammophis angolensis is a wide¬ 
spread species with several peculiar autapomorphies. It was originally described in the genus 
Amphiophis Bocage, 1872, and Brandstatter (1995) suggested that this should be revived as a 
monotypic subgenus. However, Kelly et al. (2008) did not follow this suggestion in their phyloge¬ 
netic anaylsis of Psammophis. 
Psammophis ansorgii Boulenger, 1905 
Link-Marked Sand Racer (Endemic) 
Psammophis Ansorgii Boulenger 1905:113, pi. 4, fig. 4. Holotype: BMNH 1946.1.8.49 (formerly BMNH 
1905.5.29.31) (eolleetor W.J. Ansorge). Type loeality: “between Benguella and Bihe,” Angola. 
Psammophis jallae: Loveridge (1940:62). 
Psammophis ansorgii'. Hellmieh (1957b:69), Broadley (1977b:26, 2002a:98), Brandstatter (1996:42), Wallaeh 
et al. (2014:576). ii^e 2 o°e 24 "e 
Global conservation status (lUCN): Not 
Evaluated. 
Global distribution: The species is 
known from Angola. 
Ocurrences in Angola (Map 308): The 
species is endemic to the central highlands of 
Angola. Huambo: “Bela-Vista” [-12.36667, 
16.20000] (Hellmieh 1957b:69). Undeter¬ 
mined locality: “Between Benguella to Bihe” 
(Boulenger 1905:113; Loveridge 1940:62; 
Broadley 1977b:26, 2002a:98; Wallaeh et al. 
2014:576). 
Taxonomic and distributional notes: 
Loveridge (1940) placed Psammophis ansorgii 
in the synonymy of Psammophis jallae Perac- 
ca, 1896, but Broadley (1977b) considered it a 
full species, as have subsequent authors. 
Map 308. Distribution of Psammophis ansorgii in 
Angola. 
Psammophis jallae Peracca, 1896 Jalla’s Sand Snake 
Psammophis jallae Peracca 1896:2, 2 figs. Syntypes: MZUT 1633 and 1871 (formely MZUT 1823a-b) 
(collector L. Jalla). Type locality: “Strada da Kazungula a Buluwaio” (Peracca 1896:2), [= road from 
Kazungula, Zambia to Bulawayo], Zimbabwe. 
Psammophis Rohani Angel (1922:116). Type: MNHP 20-198 (collector J. F. de Rohan-Chabot). Type locali¬ 
ty: “Region de la riviere Lumuna, affluent de Loengoue” [= Lumuna River, tributary of the Luina and 
Quando Rivers], Angola. 
Psammophis rohani'. Angel (1923:166), Monard (1937b: 128). 
Psammophis jallae'. Loveridge (1940:62), FitzSimons (1962:237), Brandstatter (1996:65), Broadley 
(1977b: 12, 1990:138, 2002a:93), Branch (1998:90), Bates et al. (2014:376), Wallaeh et al. (2014:576). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known from central and southeastern Angola, western 
Zambia, northeastern Namibia, Botswana, western and central Zimbabwe and Limpopo Province 
of South Africa. Also from the southern portion of the former Katanga Province, Democratic 
Republic of Congo {fide Wallaeh et al. 2014). 
Ocurrences in Angola (Map 309): The species occurs in central and southeastern Angola. 
Bie: “Bigondo” [-12.06667, 17.41667] (Loveridge 1940:64); “Benguella to Bihe” (Loveridge 
