MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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1940:64). Cuando Cubango: “Lumuna 
regions, Loengoue affluent” (Angel 1921:118; 
1923:168; Monard 1937b:128; Loveridge 
1940:62; Broadley 1977b: 12, 2002a:93). 
Taxonomic and distributional notes: 
Angel (1923) provided three views of the head 
(figs. 10-12) of the speeies he deseribed the 
year previously. The date of deseription has 
been given as 1921, but Angel (1923) explieit- 
ly stated that the assoeiated issue of the Bul¬ 
letin de la Soeiete Zoologique de Franee had 
appeared only in Mareh 1922. Psammophis 
rohani was synonymized with P. jallae Perraea, 
1896 by Loveridge (1940). 
Psammophis leopardinus (Bocage, 1887) 
Psammophis sibilans van leopardinus Bocage 1887a:206. Lecototype: MBL 1798 (collector J.A. d’Anchieta), 
designated hy Broadley (1977b: 18), destroyed by fire 18 March 1978. Type locality: “Catumbella” 
(Bocage 1887b:206) [= Catumbela] Benguela Province, Angola. 
Psammophis sibilans van C leopardina: Bocage (1895a: 117). 
Psammophis sibilans van E brevirostris: Bocage (1895a: 118). 
Psammophis brevirostris: Boulenger (1915:213); Monard (1937b: 133), Themido (1941:10). 
Psammophis brevirostris brevirostris: Brandstatter (1996:45). 
Psammophis sibilans sibilans: Loveridge (1957:279). 
Psammophis sibilans leopardinus: Broadley (1977b: 18). 
Psammophis brevirostris leopardinus: Brandstatter (1996:48), Branch (1998:91). 
Psammophis leopardinus: Broadley (1990:143, 2002a:95), Kelly et al. (2008:1048), Hughes and Wade 
(2002:75), Bates et al. (2014:374), Wallach et al. (2014:577). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The speeies is known from dry savanna and semi-desert regions in 
southwestern Angola and central and northern Namibia. 
Ocurrences in Angola (Map 310): This species occurs chiefly in southwestern Angola. 
Luanda: “Luanda” [-8.83333, 13.26667] (Broadley 2002:111; Hughes and Wade 2002:77). 
Huambo: “Bella Vista (Missao di Dondi)” [-12.36667, 16.20000] (Broadley 2002:111; Hughes 
and Wade 2002:77); “Huambo” [-12.76667, 15.73333] (Themido 1941:10). Benguela: “Lobito 
bay” [-12.35000, 13.5500] (Broadley 1977b:19, 2002a:lll; Hughes and Wade 2002:77); “Catum¬ 
bella” [-12.43333, 13.55000] (Bocage 1895a:117; Loveridge 1957:279; Broadley 1990:143, 
2002a:lll; Hughes and Wade 2002:77); “Quissange” [-12.43333, 14.05000] (Bocage 1895a:117); 
“Catengue” [-13.03333, 13.73333] (Broadley 2002:111; Hughes and Wade 2002:77). Huila: 
“Caconda” [-13.73333, 15.06667] (Broadley 2002:111; Hughes and Wade 2002:77); “Kalukembe” 
[-13.78333, 14.68333] (Monard 1937b:133; Broadley 2002:111); “Quillengues” [-14.06667, 
15.08333] (Bocage 1895a:114); “Vila da Ponte” [-14.46667, 16.30000] (Monard 1937b:133; 
Broadley 2002:111); “Capelongo” [-14.88333, 15.08333] (Hughes and Wade 2002:77). Namibe: 
“Capangombe” [-15.10000, 13.15000] (Bocage 1887b:206; Broadley 2002:111); “Mossamedes” 
I2"E 16.°E 2«°E 
Map 309. Distribution of Psammophis jallae in Angola. 
Leopard Sand Snake 
