350 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Loveridge 1940:30); “Mupanda” [-17.13333, 15.76667] (Monard I937b:I3I; Loveridge 1940:30); 
“Tala Kilau, Mossamedes” (Broadley 2002:113). Cuando Cubango: “Kakindo” [-15.45000, 
17.05000] (Monard 1937b:131; Loveridge 1940:30); “Kayundo” [-15.70000, 17.45000] 
(Loveridge 1940:30); “vineity of Cuito Cuanavale - approximatley 75 km W of Mavinga” 
[-15.78333, 19.70000] (Branch and McCartney 1992:2); “Cubango basin (48)” [-14.68136, 
17.44530] (Conradie et al. 2016:9, 12, 22). Undetermined Locality: “without precise location” 
(Laurent 1954:59, 1964a: 113); “Cascalada 6 km NW” (Broadley 2002:113); “Marimo 10 km W of 
Machado” (Broadley 2002:113). 
Taxonomic and distributional notes: The taxonomy of the Psammophis sibilans complex, to 
which this species belongs, remains confused (Kelly et al. 2008; Wallach et al. 2014) and we have 
here treated Angolan specimens previously refered to P. sibilans (Linnaeus, 1758) (except those 
attributed to P. zambiensis) and P. phillipsii (Hallowell, “1844” 1845) as P mossambicus (e.g., 
Loveridge 1953; Broadley 1977b; Broadley et al. 2003). Brandstatter (1996) and Hughes (1999) 
considered P phillipsii to be restricted to West Africa, and Branch (1998) was the first to use the 
name P. mossambicus Peters, 1882 to refer to the former southern and East African populations of 
P. phillipsii. Broadley (2002a) likewise recognized that the Angolan material initially assigned to 
P. sibilans and P. phillipsii should be attributed to P. mossambicus. 
Psammophis namibensis Broadley, 1975 Namib Sand Snake 
Psammophis leightoni namibensis Broadley 1975b:9, pi. 1. Holotype: TM 37093 (collector W.D. Haacke). 
Type locality: “Hams in the Uri-Hauchab Mountains, Diamond Area No. 2, South West Africa” [= Karas 
Region], Namibia. 
Psammophis leightoni namibensis: Broadley (1977b: 11, 1990:136), Brandstatter (1996:68), Branch 
(1998:90). 
Psammophis namibensis: Broadley (2002:91), Bates et al. (2014:378), Wallach et al. (2014:577). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known 
scrubland, occurring along the southwest Africa, 
Namib Desert into western Namibia, to 
Namaqualand, in the Northern Cape Province, 
South Africa. 
Ocurrences in Angola (Map 312): The 
species occurs in the southwestern Angola in 
the Namib Desert. Namibe: “Mossamedes” 
[-15.20000, 12.15000] (Broadley 1975b:ll; 
2002a: 107); “Pico Acezevedo” [-15.55000, 
12.51667] (Broadley 2002:107); “Rio Coroca” 
[-15.78333, 12.06667] (Broadley 1975b:ll; 
2002a:107); “Foz do Cunene” [-17.28333, 
11.80000] (Broadley 1975b: 11; 2002a: 107); 
“Cunene Forde, 15 km NE, Iona Res.” 
(Broadley 2002:107). 
Taxonomic and distributional notes: 
Broadley (1975b) initially described this form 
as a new subspecies of Psammophis leightoni 
Boulenger, 1902, but later considered it a full 
species (Broadley 2002). Kelly et al. (2008), 
from arid regions, including desert and succulent 
from southwestern Angola extending through the 
II^E Ili'E 20^E 24'*E 
Map 312. Distribution of Psammophis namibensis in 
Angola. 
