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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
restricted to “Tete, Mogambique” by Broadley (1966b). 
Psammophis bocagii Boulenger (1895a:538). Syntype: BMNH 67.7.23.22 (collector Donaldson Smith). Type 
locality: “Angola” later given as “Benguella,” Angola (Boulenger 1896:161). 
Psammophis sibilans var. A (var. subtaeniata): Bocage (1895a: 116). 
Psammophis bocagii: Boulenger (1896:161, 1915:213), Monard (1937b: 131), Bogert (1940:82). 
Psammophis Bocagii: Bocage (1897a:201). 
Psammophis subtaeniatus subtaeniatus: Loveridge (1940:55), Broadley (1966b:6, 1977b: 13), Brandstatter 
(1996:99). 
Psammophis subtaeniatus: Branch (1998:91), Broadley (1990:342, 2002b:93), Bates et al. (2014:379), 
Wallach et al. (2014:580), Conradie et al. (2016:22). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from southern Angola and northern Namibia, east 
through Botswana to southern Zambia, Zimbabwe, parts of western Mozambique, the northeastern 
provinces of South Africa and eastern Swaziland. 
Ocurrences in Angola (Map 314): The 
species occurs in southern Angola. Benguela: 
“Catumbella” [-12.43333, 13.55000] (Bocage 
1895a:115, 1897a:201; Loveridge 1940:55; 
Broadley 2002:108); “Benguella” [-12.58333, 
13.41667] (Boulenger 1896:161; Loveridge 
1940:55; Broadley 1966b:8, 1977b: 13, 
2002a:108). Huila: “Molundo” [-15.01667, 
13.20000] (Monard 1937b:131; Loveridge 
1940:55). Namibe: “Biballa” [-14.76667, 
13.36667] (Bocage 1895a:115, 1897a:201; 
Loveridge 1940:55); “Chao de Chella” 
[-14.88944, 13.27417] (Broadley 2002:108); 
“Maconjo” [-15.01667, 13.20000] (Bocage 
1895a:115, 1897a:201; Loveridge 1940:55; 
Broadley 2002:108). Cunene: “Mupa” 
[-16.18333, 15.75000] (Bocage 1897a:201; 
Loveridge 1940:55; Broadley 2002:108); 
“Cahama 5 km SE” [-16.28333, 14.30000] 
(Broadley 2002:108); “Humbe” [-16.68333, 14.90000] (Bocage 1895a:115, 1897a:201; Loveridge 
1940:55; Broadley 2002:108); “Forte Roqadas” [-16.71667, 15.01667] (Monard 1937b:131; 
Loveridge 1940:55; Broadley 2002:108); “Dongwenna, Mossamedes” [-17.01667, 14.71667] 
(Broadley 2002:108). “Mupanda” [-17.13333, 15.76667] (Monard 1937b:131; Loveridge 1940:55; 
Broadley 2002:108); “Cunene” (Bocage 1895a: 114; Loveridge 1940:55; Broadley 2002:108). 
Cuando Cubango: “Cuito drainage” [-17.57333, 23.26000] (Conradie et al. 2016:9-10, 22). 
Undetermined Locality: “Cuanza” (Bocage 1895a: 115); “Rio Bengo” (Bocage 1895a: 115, 
1897a:201; Loveridge 1940:55; Broadley 2002:108); “Pen Pen” (Broadley 2002:108). 
Taxonomic and distributional notes: Psammophis subtaeniatus has been known by a variety 
of names, Boulenger (1895a) restricted the name subtaeniatus to the eastern form and erected a 
new name, Psammophis bocagii (Boulenger 1895), for the western race based on specimens from 
Angola (Broadley 1977b). The description appears as part of a key and no precise locality or num¬ 
ber of specimens is given, however, the fact that a range of scale counts is provided demonstrates 
that there was originally a syntype series. Boulenger (1896) indicated that the single BMNH spec¬ 
imen had come from Bocage in Lisbon, so it is likely that at least one syntype remained in the MBL 
Map 314. Distribution of Psammophis subtaeniatus in 
Angola. 
