MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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collection, perhaps not marked as such. Bocage (1895a) referred to P. sibilans var. A, but in 
the same aceount equated this with P. s. var. subtaeniatus Peters, 1882. Loveridge (1940) used 
P subtaeniatus subtaeniatus for the western form and placed P. bocagii in its synonymy. 
Psammophis trigrammus Gunther, 1865 Western Sand Snake 
Psammophis trigrammus Gunther 1865b:95, pi. 2, fig. e. Holotype: BMNH 1946.1.8.12 (formerly BMNH 
64.7.11.4) (eolleetor J. Monteiro). Type loeality: “on the ba nks of river S. Nieolao (Little Fish Bay, West 
Afriea)” [= Rio Sao Nieolau], Namibe Provinee, Angola. 
Psammophis trigrammus: Boeage (1887b:206), Loveridge (1940:23), FitzSimons (1962:221), Broadley 
(1977b:9, 1990:133, 2002a:87), Brandstatter (1996:104), Braneh (1998:89), Bates et al. (2014:379), 
Wallaeh et al. (2014:581). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The speeies is known from southern Angola and western Namibia, 
reaching its southern limit in the extreme north of the Richtersveld, South Africa. 
Ocurrences in Angola (Map 315): The 
species is restrieted to Namibe Province. 
Namibe: “Catara River” [-13.55000, 
12.55000] (Broadley 2002:103); “on the banks 
of river S. Nieolao (Little Fish bay. West Afri- 
ka)” [-14.26667, 12.36667] (Gunther 
1865b;95); “Rio de Sao Nieolau, Mossamedes 
(Rio San Nieolau)” [-14.26667, 12.36667] 
(Bocage 1887b:206; Loveridge 1940:23; 
FitzSimons 1962:221; Broadley 1977b:9, 
1990:133, 2002ba:86, 103; Wallaeh et al. 
2014:581); “Rio Coroca” [-15.78333, 
12.06667] (Broadley 2002:103); “Iona 
Reserve, 7 km to Oncocua” [-16.85831, 
12.61275] (Broadley 2002:103). 
Taxonomic and distributional notes: 
Bocage (1887b) corrected the type locality to 
“Mossamedes” [= Rio Sao Nieolau, Namibe]. 
20°E 24'E 
Map 315. Distribution of Psammophis trigrammus in 
Angola. 
Psammophis zambiensis Hughes and Wade, 2002 Zambian Whip Snake 
Psammophis zambiensis Hughes and Wade 2002:75. Holotype: BMNH 1959.1.1.81 (eolleetor H.J. Bredo). 
Type loeality: “supposedly from “Abereom” (= Mbala) area of Zambia ... but likely to be from Mwem- 
Wantipa”. 
Psammophis sibilans sibilans: Laurent (1950a:9, 1954a:59), Thys van den Audenaerde (1966:34). 
Philothamnus irregularis irregularis: Managas (1973:191). 
Psammophis zambiensis: Broadley and Cotterill (2004:49), Wallaeh et al. (2014:582). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The speeies is known with certainty from the southern Demoeratic 
Republic of Congo and northern Zambia. Literature records of Psammophis sibilans that may be 
referable to P. zambiensis exist from the Caprivi Strip of Namibia and adjacent northern Botswana, 
northern Zimbabwe, Malawi, Zambezia Province of Mozambique and northestem Angola. 
Ocurrences in Angola (Map 316): None eonfirmed but Psammophis zambiensis is expected 
to occur in northeastern regions of the country. The following records are P. ^sibilans" records that 
