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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
may be referable to P. zambiensis. Mexico: 
“Calombe” [-11.83333, 19.93333] (Managas 
1973:191). Lunda Norte: “Dundo” [-7.36667, 
20.83333] (Laurent 1950a:9, 1954a:59, 
1964a: 113; Thys van den Audenaerde 
1966:34); “Barrage de la Luachimo” 
[-7.38333, 20.85000] (Thys van den Audenaer¬ 
de 1966:34); “Dundo, R. Mussungue, aff. 
Luachimo” [-7.41667, 20.83333] (Thys van 
den Audenaerde 1966:34); “Muita (Luembe 
E)” [-7.80000, 21.45000] (Laurent 1954a:59). 
Taxonomic and distributional notes: 
Hughes and Wade (2002) found significant dif¬ 
ferences between specimens of '‘Heopardinus^^ 
from Angola and from Zambia and described 
P. zambiensis to accommodate the latter. Some 
eastern records of P. leopardinus could corre¬ 
spond to this new species. Hughes and Wade 
(2002) presented information from Desmond 
Vesey-FitzGerald via Donald G. Broadley that substantiated the likely origin of the holotype from 
Mweru-Wantipa in the Mporokoso District [Zambia]. Hughes and Wade (2002) plotted localities 
of Psammophis 'sibilans' from the literature, which could include material assignable to P. zambi¬ 
ensis. Two such Angolan records correspond to literature references from Laurent (1950, 1954a, 
1964a) and Thys van den Audenaerde (1966). Wallach et al. (2014) included only the former 
province of Katanga (Democratic Republic of Congo) and northern Zambia in the distribution of 
the species, corresponding to the verified records given in the description. 
Genus Psammophylax Fitzinger, 1843 
Psammophylax acutus (Gunther, 1888) Striped Beaked Snake 
Psammophis acutus Gunther 1888:327, pi. 19, fig. d. Holotype: BMNH 1946.1.2.81 (formerly BMNH 1864. 
7.13.36) (collector F.M.J. Welwitsch). Type locality: “Pungo Andongo,” Malanje Pronvince, Angola. 
Psammophis oxyrhynchus: Gunther (1865a:480, 1895:89). 
Rhagerhis acuta: Bocage (1895a:111). 
Rhamphiophis acutus: Boulenger (1896:148, 1905:113, 1915:212), Loveridge (1933:252), Hellmich 
(1957b:71), Managas (1973:195), Broadley et al. (2003), Spawls et al. (2004:398), Chippaux (2006:170). 
Rhamphiophis acutus {Rhagerhis acuta): Monard (1937h:128). 
Rhamphiophis acutus wittei: Laurent (1964a:111). 
Rhamphiophis acutus acutus: Loveridge (1957:277), Laurent (1964a:111), Broadley (1971h:3), Chirio and 
Ineich (1991:220). 
Rhamphiophis acutus jappi: Broadley (1971d:4). 
Psammophylax tritaeniatus tritaeniatus: Mananas (1973:194). 
Psammophylax acutus acutus: Kelly et al. (2008:1052). 
Psammophylax acutus jappi: Kelly et al. (2008:1052). 
Psammophis acutus: Wallach et al. (2014:575). 
Psammophylax acutus: Conradie et al. (2016:22). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is known from Gabon to the former province of Katanga 
(Democratic Republic of Congo) to Angola and east to northern Malawi, western Tanzania and 
ll-E 20°E ,24'’E 
Map 316 Distribution of Psammophis zambiensis in 
Angola. 
