MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
355 
Burundi. Records from northern Namibia 
(M. Griffin in Wallach et al. 2014) have not 
been verified or vouchered. 
Ocurrences in Angola (Map 317): The 
species occurs from southwestern regions of 
Huila Province to central and northeastern 
Angola. Lunda Norte: “Dundo” [-7.36667, 
20.83333] (Laurent 1964a:lll; Broadley 
1971d:4); “Cassange” [-9.58333, 17.86667] 
(Bocage 1895a:lll; Broadley 1971d;3). 
Lunda Sul: “Alto Cuilo” [-10.01667, 
19.55000] (Laurent 1964a;111; Broadley 
1971d:3); “Alto Chicapa” [-10.88333, 
19.23333] (Laurent 1964a:lll; Broadley 
1971b:3). Moxico: “Calombe, Luso” 
[-11.83333, 19.93333] (Managas 1973:194, 
195). Malanje: “Pungo-Andongo” [-9.66667, 
15.58333] (Gunther 1865a:480; 1888:327, 3*’' Distribution of Psammophylax acums in 
1895:89; Bocage 1895a: 111; Boulenger 
1896:148; Loveridge 1933:252; Monard 1937b; 128; Broadley 1971d:3; Chirio and Ineich 
1991:220; Chippaux 2006:170; Wallach et al. 2014:575). Huambo: “Bela-Vista” [-12.36667, 
16.20000] (Hellmich 1957b:71; Broadley 1971d:3). Benguela: “Bigondo” [-12.06667, 17.41667] 
(Monard 1937b:128). Huila: “Caconda” [-13.73333, 15.06667] (Bocage 1895a:lll; Monard 
1937b:128; Broadley 1971d:3); “Huilla” [-15.05000, 13.55000] (Bocage 1895a:lll; Monard 
1937b:128; Broadley 1971d:3). Cuando Cubango: “Longa River (23)” [-14.58972, 18.17083] 
(Conradie et al. 2016:8-9, 22); “Longa River (51)” [-14.58970, 18.1711] (Conradie et al. 2016:8, 
12, 22). Undetermined locality: “Between Benguella and Bihe” (Boulenger 1905:113; Broadley 
1971d:3). 
Taxonomic and distributional notes: Broadley (1971b) recognized three subspecies: 
Rhamphiophis a. acutus, R. a.jappi (Broadley, 1971) andR «. togoensis (Matschie, 1893) and pro¬ 
visionally synonymized Rhamphiophis acutus wittei (Laurent, 1956) with the nominate form. 
Recently Kelly et al. (2008) showed that Rhamphiophis is diphyletic and transfered R. acutus and 
its subspecies to the genus Psammophylax. The specimen from “Dundo” (MD 5193-A) identified 
by Laurent (1964a) as R. a. acutus and later as R. a.jappi by Broadley (197 Id) is here interpreted 
as the typical form. We recently examined the specimen identified by Mananas (1973) as Psam¬ 
mophylax tritaeniatus tritaeniatus (Gunther, 1868) from “Calombe, Luso” deposited in the Institu¬ 
te de Investigagao Cientifica Tropical, Lisboa and it is likewise referable to Psammophylax a. acu¬ 
tus. Wallach et al. (2014) erroneously allocated R. acutus to Psammophis. 
Psammophylax rhombeatus ocellatus (Bocage, 1873) Spotted Skaapsteker (Endemic) 
Psammophylax ocellatus Bocage 1873a:22L Holotype: MBL specimen number unknown (collector J.A. 
d’Anchieta), specimen lost fide Broadley (1977c). Type locality: “I’interieur de Mossamedes (Gambos)” 
[= Gambos], Huila Province, Angola. 
Psammophis rhombeatusl: Bocage (1867b:224). 
Psammophylax rhombeatus: Bocage (1895a: 108), Boulenger (1896:138, 1915:211), Bates et al. (2014), 
Wallach et al. (2014:582). 
Trimerorhinus rhombeatus: Monard (1937b: 126, 128). 
Psammophylax rhombeatus oeellatus: Broadley (1977c:20, 1990:121), Branch (1998:88). 
