370 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
west Angola. A subadult Naja annulata from Lunda Norte Province (Branch and Conradie 2015), 
is the first voucheredd record for the country. 
Naja melanoleuca Hallowell, 1857 Forest Cobra 
Naja haje (var. melanoleuca) Hallowell 1857:61. Syntypes: ANSP 6875-78 (collector H. A. Ford). Type loca¬ 
lity: “Gaboon country, West Africa” [= Gabon]. 
Naja haje\ Bocage (1866a:51, 1895a:132), Peters (1877a:618). 
Naja haje haje: Loveridge (1957:291). 
Naja melanoleuca: Boettger (1898:120), Boulenger (1905:114, 1915:219), Ferreira (1900b:133, 1903:12), 
Parker (1936:126), Bogert (1940:87), Laurent (1950a: 10, 1954a:60), Hellmich (1957b:72), FitzSimons 
(1962:300), Broadley (1968c:5), Mananas (1981:27), Broadley (1990:291), Spawls and Branch 
(1995:71), Branch (1998:107), Broadley and Cotterill (2004:47), Chippaux (2006:225), Trape and Mane 
(2006:198), Dobiey and Vogel (2007:69), Bates et al. (2014:404), Wallach et al. (2014:120), Ceriaco et 
al. (2017:141). 
Naja melanoleuca melanoleuca: Laurent (1964a: 120), Thys van den Audenaerde (1966:35), Chirio and 
LeBreton (2007:582). 
Naja (Boulengerina) melanoleuca: Wallach et al. (2009:31). 
Boulengerina melanoleuca: Wallach et al. (2014:120). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The N. melanoleuca species complex is known from forest and savanna 
through west, central and east Africa, from Senegal east to western Ethiopia, southern Somalia, 
south to Angola on the west coast and KwaZulu-Natal, South Africa on the east coast. Populations 
in the east of the range are partly disjunct. With the recent elevation of N. subfulva to specific 
status (Ceriaco et al. 2017), the global distribution of V. melanoleuca will need to be reevaluated. 
Ocurrences in Angola (Map 333): The 
species occurs from the northeast of the coun¬ 
try, including areas adjacent with Zambia in 
Mexico Province (pers. obs.) to the northwest, 
including Cabinda, and throughout western 
Angola, exclusive of arid regions. Cabinda: 
“Chinchoxo” [-5.10000, 12.10000] (Bocage 
1866a:51; Ferreira 1900b:133; Mananas 
1981:27); “Cabinda” [-5.55000, 12.18333] 
(Peters 1877a:618; Bocage 1895a: 132, Boett¬ 
ger 1898:120). Lunda Norte: “Dundo” 
[-7.36667, 20.83333] (Laurent 1950a: 10, 
1954a;60, 1964a; 120; Thys van den Audenaer¬ 
de 1966:35; Mananas 1981:27); “Dundo, R. 
Capemba” (Thys van den Audenaerde 
1966:35); “Dundo, Barrage de la Luachimo” 
[-7.38333, 20.85000] (Thys van den Audenaer¬ 
de 1966:35); “Dundo, Cacanda” [-7.40000, Map 333. Distribution ofiVayame/anoto in Angola. 
21.80000] (Thys van den Audenaerde 1966:35); “Cossa” [-7.93333, 21.38333] (Thys van den 
Audenaerde 1966:35); “R. Camaiala” [-8.05000, 18.61667] (Thys van den Audenaerde 1966:35). 
Malanje: “Duque de Braganga” [-9.10000, 15.95000] (Bocage 1866a;51, 1895a:132; Ferreira 
1900b:133; Mananas 1981:27); “Pungo-Andongo” [-9.66667, 15.58333] (Boulenger 1905:114; 
Mananas 1981:27). Kwanza Norte: “Piri-Dembos” [-8.56667, 14.50000] (Hellmich 1957b:72; 
Mananas 1981:27; Ceriaco et al. 2017:141); “Ambaca” [-9.26667, 15.18333] (Mananas 1981:27); 
