MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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“N’dalla Tando” [-9.30000, 14.91667] (Ferreira 1903:12; Mananas 1981:27); “Cazengo” 
[-9.33333, 14.76667] (Mana 9 as 1981:27). Kwanza Sul: “Libolo-Luati” [-9.98333, 14.90000] 
(Hellmieh I957b:72; Managas 1981:27; Ceriaeo et al. 2017); “Quirimbo” [-10.68333, 14.26667] 
(Parker 1936:126; Managas 1981:27); “Congulu” [-10.86667, 14.28333] (Parker 1936:126; Mana- 
gas 1981:27). Huambo: “Galanga” [-12.06667, 15.15000] (Boeage I895a:I32). Benguela: “Caha- 
ta” [-12.35000, 14.81667] (Boeage I895a:I32; Ferreira I900b:I33; Managas 1981:27); “Hanha” 
[-12.25000, 13.75000] (Bogert 1940:87; Mananas 1981:27). Hufla: “Sanguengue” [-12.36667, 
16.20000] (Hellmieh 1957b:72); “Huilla” [-15.05000, 13.55000] (Ferreira 1900b:133; Mananas 
1981:27). Namibe: “Capangombe” [-15.10000, 13.15000] (Ferreira 1900b:133; Mananas 
1981:27). 
Taxonomic and distributional notes: Broadley and Cotterill (2004) referred to Naja 
melanoleuca as a “species complex” that might contain several cryptic species. It was being 
reviewed by Broadley et al. at the time of the senior author’s death. Wallach et al. (2009) assigned 
Naja melanoleuca to the subgenus Boulengerina and later (Wallach et al. 2014) treated Boulenge- 
rina as generically distinct. Although Spawls and Branch (1995) and Dobiey and Vogel (2007) 
considered N. melanoleuca to be restricted to northern Angola, including the Cabinda enclave, it is 
much more widely spread throughout the country. Ceriaeo et al. (2017) presented molecular evi¬ 
dence to support Naja subfulva Laurent, 1955, as a valid species and not a mere “savannah form” 
or subspecies of N. melanoleuca as considered by many previous authors. The two species are eas¬ 
ily told apart by a combination of morphological characters and coloration, but to separate the 
Angolan records, many of them from before the original description of subfulva, will require a 
detailed review of all available specimens. Despite N. melanoleuca being considered more of a 
forest dweller and subfulva a savannah species, they are sympatric in parts of Angola (LMPC pers. 
obs.), frustrating attempts to infer the identity of records based solely on distribution. 
Naja mossambica Peters, 1854 Mozambique Spitting Cobra 
N[aja]. mossambica Peters 1854:625. Leetotype: ZMB 2811 (eolleetor W.C.H. Peters), designated by Golay 
et al. (1993:188). Type loeality: “Tette and Sena” [= Tete, Vila de Sena], Mozambique, restrieted to “Tette” 
as a result of leetotype designation. 
Naja mossambica mossambica: Managas (1981:28). 
Naja mossambica: Broadley (1966e:25, 1974:156), Broadley (1990:295), Spawls and Braneh (1995:74), 
Broadley and Cotterill (2004:47), Dobiey and Vogel (2007:71), Wurster et al. (2007:438) 
Naja mossambica mossambica: Managas (1981:28). 
Naja (Afronaja) mossambica: Wallaeh et al. (2009:32), Conradie et al. (2016:22). 
Afronaja mossambica: Wallach et al. (2014:10). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The species is widespread in eastern and southern Africa, occuring from 
southern Tanzania southwestwards to extreme southeastern Angola, northern Namibia and 
Botswana, and southwards to eastern South Africa and Swaziland. 
Ocurrences in Angola (Map 334): The species occurs in the extreme southeastern regions of 
Cuando Cubango Province and there is an isolated record in Namibe Province. Namibe: “Macon- 
jo” [-15.01667, 13.20000] (Broadley 1974:156; Managas 1981:28). Cuando Cubango: “Cuito 
basin (30a)” [-17.50875, 20.06594] (Conradie et al. 2016:9, 10, 22); “Cuando basin (44b)” 
[-17.56916, 23.27305] (Conradie et al. 2016:9, 10, 22). 
Taxonomic and distributional notes: Dobiey and Vogel’s (2007) distribution map shows that 
the Mozambique Spitting Cobra in Angola is restricted to the extreme southeastern regions of 
Cuando Cubango Province, near the Zambian and Namibian borders. However, Broadley (1974, 
