MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Naja mossambica nigricincta: FitzSimons (1962:306), Managas (1981:28). 
Naja nigricincta'. Broadley (1966c:25), Wurster et al. (2007:445). 
Naja nigricollis nigricincta'. Broadley (1968c:7), Spawls and Branch (1995:78). 
Naja nigricincta nigricincta'. Broadley (1974:158, 1990:294), Dobiey and Vogel (2007:72). 
Naja (Afronaja) nigricincta'. Wallach et al. (2009:32). 
Afronaja nigricincta'. Wallach et al. (2014:10). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The nominate form of the speeies is known from southwestern Angola 
and eentral and northern Namibia. Naja nigricincta woodi Pringle, 1955 is extralimital, oeeurring 
in southern Namibia and western South Africa. 
Ocurrences in Angola (Map 335): The 
species is known from southwestern Angola. 
Benguela: “Benguela” [-12.58333, 13.41667] 
(Bocage 1866a:51, 1866b:71, 1867b:228, 
1895a:136; Monard 1937b;136; Broadley 
1974:158; Managas 1981:28); “Rio Equimina” 
[-13.20000, 14.78333] (Broadley 1974:158; 
Mananas 1981:28). Namibe: “Munhino” 
[-14.91667, 13.00000] (Bogert 1940:89; Fitz¬ 
Simons 1962:306; Broadley 1968c:7, 
1974:158, 1990:294; Managas 1981:28; Wal¬ 
lach et al. 2014:10); “Maconjo” [-15.01667, 
13.20000] (Broadley 1974:158; Managas 
1981:28); “Capangombe” [-15.10000, 
13.15000] (Bocage 1895a:136); “Foz do Cune- 
ne” [-17.28333, 11.80000] (Broadley 
1974:158; Managas 1981:28). 
™ . j j* X M X* 1 X Map 335. Distribution of Vo/a «/er/cmcto in Angola. 
Taxonomic and distributional notes: 
Bocage (1895a) recognized several varieties of Naja nigricollis Reinhardt, 1843 based on 
coloration (var. occidentalis; var. melanoleuca', Yar.fasciata). Naja nigricollis nigricinctus Bogert, 
1940 is a synonym of Bocage’s (1895a) Yav.fasciata, but since the older name was preoccupied by 
Najafasciata Laurenti (1768), a synonym ofNaja naja (Linnaeus, 1758), Broadley (1966c) accept¬ 
ed Bogert’s (1940) as the valid name of this species, treating it as a subspecies of N. nigricollis. 
Years later he treated this form as a subspecies of A. mossambica Peters, 1854 (Broadley 1968c, 
1974). Bocage had recorded A. n. y 2 lv. fasciata from “Dondo,” however Broadley (1974) examined 
the Museu Bocage specimens and only found one specimen from that locality, which he nominat¬ 
ed as the lectotype of Naja nigricollis var. occidentalis (Bocage 1895a) (= Naja nigricollis Rein¬ 
hardt, 1843) whereas the A. nigricollis var. fasciata speciemens were from “Benguela” and 
“Maconjo,” Spawls and Branch (1995) still recognized nigricincta as a subspecies of nigricollis, 
whereas Wiister et al. (2007) treated it as a separate evolutionary species, with A. n. woodi as a 
subspecies. 
Naja nigricollis Reinhardt, 1843 Black-Necked Spitting Cobra 
Naja nigricollis Reinhardt 1843:269, pi. 3, figs. 5-7. Lectotype: ZMUC 65204 (collector J.R. Chenon) desig¬ 
nated by F. W. Braestmp in Hughes and Barry (1969:1027). Type locality: “Guineakysten” (Reinhardt 
1843:271) [= coast of Ghana fide Hughes and Barry (1969:1027)], probably near “Christiansborg Castle, 
now a suburb of Accra,” [Ghana], where the collector was stationed (Rasmussen and Hughes 1997:15). 
