374 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Naja nigricollis var. occidentalis Bocage 1895a: 135. Lectotype: MBL 1963 (collector F.A.P. Bayao), 
destroyed by fire 18 March 1978, designated by Broadley (1974:156). Type locality: “Dondo ... Quis- 
sange, Quillengues, Huilla et Humbe [Angola] ... Bissau [Guinea-Bissau],” restricted to “Dondo” 
[= Sumbe] Kwanza Norte Province, Angola through the designation of the lectotype. 
Naja nigricollis var. melanoleuca Bocage 1895a: 136 (non Hallowell, 1857). Lectotype: MBL 1972 (collector 
J.A. d’Anchieta), destroyed by fire 18 March 1978, designated by Broadley (1974:156). Type locality: 
“Catumbella et... Caconda” restricted to “Caconda,” Angola through the designation of the lectotype. 
Naja nigricollis-. Bocage (1867b:228, 1896a:113), Peters (1881:149), Boettger (1898:120), Ferreria 
(1900b: 134), Boulenger (1905:114), Schmidt (1933:14), Monard (1937b: 136-137), Broadley (1966c:25), 
Mananas (1981:28), Spawls and Branch (1995:78), Dobiey and Vogel (2007:74), Wiirst et al. (2007:438). 
Naia nigricollis-. Boulenger (1896:378). 
Naja nigricollis nigricollis-. Mertens (1938a:442), Laurent (1950a: 10, 1954a:60), Hellmich (1957a:74, 
1957b:73), Loveridge (1957:292), FitzSimons (1962:302), Broadley (1974:156, 1990:292). 
Naja nigricollis occidentalis-. Laurent (1964a: 119), Thys van den Audenaerde (1966:35), Broadley (1968c:7). 
Naja (Afronaja) nigricollis-. Wallach et al. (2009:32). 
Afronaja nigricollis-. Wallach et al (2014:10). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution; The speeies is widely distributed from southern Mauritania and Senegal 
east to Ethiopia and Somalia, and south to Zambia, Malawi, the southern Democratie Republic of 
Congo, and Angola. Absent from the forests of Central and West Africa, with disjunct populations 
along the lower Congo River, in southwestern Angola and elsewhere. 
Ocurrences in Angola (Map 336): The 
species very widespread, from coastal areas to 
the interior highlands of Angola. Zaire: 
“Ambrizette” [-7.23333, 12.86667] (Bocage 
1895a: 135; Boettger 1898:120; Mananas 
1981:28). Malanje: “Malanje” [-9.55000, 
16.35000] (Peters 1881:149; Bocage 
1895a: 135; Managas 1981:28). Kwanza 
Norte: “Golungo Alto” [-9.13333, 14.76667] 
(Boulenger 1905:114; Monard 1937b: 136); 
“Dondo” [-9.68333, 14.43333] (Bocage 
1895a: 135; Ferreira 1900b: 134; Monard 
1937b: 136; Loveridge 1957:292; Hellmich 
1957b:73; Broadley 1968c:7; Mana 9 as 
1981:28). Kwanza Sul: “Libolo-Luati” 
[-9.98333, 14.90000] (Hellmich 1957b:73; 
Managas 1981:28); “Dondo a Humbe” (Ferrei¬ 
ra 1900b:134). Bie: “Chitau” [-11.43333, 
17.15000] (Schmidt 1933:14). Benguela: 
1867b:228, 1895a:135; Ferreira 1900b:134; 
Map 336. Distribution of Naja nigricollis in Angola. 
Catumbella” [-12.43333, 13.55000] (Bocage 
Mananas 1981:28); “Quissange” [-12.43333, 
14.05000] (Bocage 1895a:135; Ferreira 1900b:134; Monard 1937b:136; Loveridge 1957:292; 
Broadley 1968c:7; Managas 1981:28); “Chivitidi/Entre Rios” [-13.01667, 14.63333] (Hellmich 
1957a:74; Managas 1981:28); “Cubal” [-13.03333, 14.25000] Mertens 1938:442; Managas 
1981:28); “Hanha” [-13.30000,14.20000] (Bocage 1896a:113; Managas 1981:28). Huila: “Cacon¬ 
da” [-13.73333, 15.06667] (Bocage 1895a:135; Ferreira 1900b:134; Monard 1937b:136; Managas 
1981:28); “Quillengues” [-14.06667, 15.08333] (Bocage 1895a:135; Monard 1937b:136; Loverid¬ 
ge 1957:292; Broadley 1968c:7; Managas 1981:28); “Kuvangu/Kubangu/Vila da Ponte” 
