MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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[-14.46667, 16.30000] (Monard I937b:I36-I37; Managas 1981:28); “Capelongo” [-14.88333, 
15.08333] (Managas 1981:28); “Huilla” [-15.05000, 13.55000] (Bocage I895a:I35; Monard 
I937b:I36; Loveridge 1957:292; Broadley I968c:7; Mana 9 as 1981:28); “Osi” [-15.08333, 
15.40000] (Monard I937b:I36-I37; Managas 1981:28); “Gambos” [-15.76667, 14.10000] (Mana- 
9 as 1981:28). Namibe: “Capangombe” [-15.10000, 13.15000] (Ferreira I900b:I34; Monard 
I937b:I36; Managas 1981:28). Cunene: “Humbe” [-16.68333, 14.90000] (Boeage I895a:I35; 
Ferreira 1900b: 134; Monard 1937b: 136; Loveridge 1957:292; Broadley I968c:7; Mana 9 as 
1981:28); “Mupanda” [-17.13333, 15.76667] (Monard I937b:I36-I37; Managas 1981:28). Cuan- 
do Cubango: “Kayundu” [-15.70000, 17.45000] (Monard I937b:I36-I37; Mananas 1981:28). 
Taxonomic and distributional notes: Bocage’s (1895a) locality of “Ambrizette” in Zaire 
Province has been considered questionable and has been excluded from some distribution maps for 
the species (Spawls and Branch 1995; Dobiey and Vogel 2007). However, Dobiey and Vogel 
(2007) do include Cabinda within the range of the species, although we are not aware of any 
published records. These authors also indicate occurrence in northcentral Namibia, although only 
a single specimen, from Katima Mulilo in the eastern Caprivi Strip has been documented from that 
country (Broadley 1974). The taxonomy of N. nigricollis still requires further study (Wiister et al. 
2007). Wallach et al. (2009) placed the African spitting cobras in a new subgenus, Afronaja 
Wallach, Wiister and Broadley, 2009, with Naja nigricollis as the type species and Wallach et al. 
(2014) have treated this as a full genus. 
Naja subfulva Laurent, 1955 Savanna Cobra 
Naja melanoleuca subfulva Laurent 1955:61. Holotype: MRAC 17514 (collector Van der Borght). Type loca¬ 
lity: “Lwiro, 1850 m, Terr, de Kabare, Kivu,” Democratic Republic of the Congo. 
Naja subfulva: Ceriaco et al. (2017:141). 
Global conservation status (lUCN): Not Evaluated. 
Global distribution: The N. melanoleuca species complex is known from forest and savanna 
through west, central, and east Africa, from Senegal east to western Ethiopia, southern Somalia, 
south to Angola on the west coast and KwaZulu-Natal, South Africa on the east coast. Populations 
in the east of the range are partly disjunct. With 
the recent elevation of N. subfulva to specific 
status (Ceriaco et al. 2017), the global distribu¬ 
tion of both this taxon and N. melanoleuca 
sensu stricto will need to be reevaluated. 
Ocurrences in Angola (Map 337): The 
species should occur across the country, except 
in the more heavily forested areas in the north, 
but currently the only confirmed records are for 
Kwanza Norte, Kwanza Sul and Malanje 
provinces. Malanje: “Cacuso” [-9.421023, 
15.746679] (Ceriaco et al. 2017:141). Kwan¬ 
za: Norte: “Piri-Dembos” [-8.56667, 
14.50000] (Ceriaco et al. 2017:141). Kwanza 
Sul: “Libolo-Luati” [-9.98333, 14.90000] 
(Ceriaco et al. 2017:141). 
Taxonomic and distributional notes: See 
Naja melanoleuca account. 
Il-E t6°E 20°E 24“ E 
Map 337. Distribution of Naja subfulva in Angola. 
