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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
“Fazenda Otilia, Encoge” [-7.55000,15.03333] 
(Managas 1973:191). Lunda Norte; “Dundo” 
[-7.36667, 20.83333] (Laurent I950a:8, 
I954a:47, 1964a: 106; Thys van den Audenaer- 
de 1966:32); “Muita (Luembe E)” [-7.80000, 
21.45000] (Laurent I954a:47). Luanda: 
“Loanda” [-8.83333, 13.26667] (Bocage 
I895a:85; Ferreira I900a;5I; Monard 
1937b: 114). Bengo: “Cacolo ao Rio Bengo” 
(Ferreira I900a;5I). Malanje: “Duque de Bra- 
gan 9 a” [-9.10000, 15.95000] (Bocage 
I866a;49, I882b:5, I895a;85; Monard 
1937b: 114); “Pungo-Andongo” [-9.66667, 
15.58333] (Gunther I865a:480; Bocage 
I882b:5, I895a:85; Ferreira 1906:167; Monard 
1937b: 114). Mexico: “Lago Dilolo” 
[-11.50000, 22.01667] (Managas 1973:191). 
Huambo: “Bela-Vista” [-12.36667, 16.20000] 
Map 353. Distribution of Philothamnus angolensis in 
Angola. 
(Hellmich 1957b:64); “Mt. Moco” [-12.41667, 
15.18333] (Parker 1936:125). Kwanza Norte: “Piri-Dembos” [-8.56667, 14.50000] (Hellmich 
1957b:64); “Ambaca” [-9.26667, 15.18333] (Ferreira 1900a:51); “Cazengo” [-9.33333, 14.76667] 
(Ferreira 1900a;51); “Cambondo” [-9.15963, 14.65771] (Boulenger 1893:356; Ferreira 1906:167, 
Monard 1937b:114, Mana 9 as 1973:193); “N’dalla Tando” [-9.30000, 14.91667] (Ferreira 
1903:10). Kwanza Sul: “Congulu” [-10.86667, 14.28333] (Parker 1936:125); “Chingo” 
[-11.20000, 13.85000] (Ferreira 1904:115; Monard 1937b:114). Bie: “Silva Porto” [-12.38333, 
16.95000] (Mana 9 as 1973:191). Benguela: “Quissange” [-12.43333, 14.05000] (Bocage 
1895a:85; Monard 1937b:114); “Quindumbo” [-12.46667, 14.93333] (Bocage 1895a:85; Monard 
1937b:114); “Alto Cubal” [-13.03333, 14.25000] (Mertens 1938a:439; Hellmich 1957b:64); 
“Hanha” [-13.30000, 14.20000] (Bocage 1896a:112). Huila: “Caconda” [-13.73333, 15.06667] 
(Boulenger 1893:96; Bocage 1895a:85; Monard 1937b:114); “Capelongo” [-14.91667, 15.08333] 
(Bogert 1940:53); “Kuvangu/Vila-da-Ponte” [-14.46667, 16.30000] (Monard 1937b: 121); “Huilla” 
[-15.05000, 13.55000] (Bocage 1895a:85, 1887b:205, 1895a:85; Monard 1937b:114). Namibe: 
“Capangombe” [-15.10000, 13.15000] (Bocage 1882b:6, 7, 1897a:200; Loveridge 1951:9, 
1957:261; Broadley 1990:238; Chippaux 2006:135; Chirio and LeBreton 2007:500; Wallach et al. 
2014:52); “Mossamedes” [-15.20000, 12.15000] (Bocage 1887b:205, 1895a:85). Cunene: “Ruis- 
seauMbale” [-15.16667,16.75000] (Monard 1937b:121); “Mupa” [-16.18333,15.75000] (Monard 
1937b:121). Undetermined Locality: “Cuango” (Peters 1881:149) (Malanje Province impossible 
to georeference; See History Section for more detailed information). 
Taxonomic and distributional notes: The majority of Angolan literature records for 
Philothamnus irregularis (Leach, 1819) are probably referable to P. angolensis or P. hoplogaster, 
two species with which it has commonly been confused (Hughes 1985). Philothamnus irregularis 
is essentially a West Africa form ranging from Senegal to northern Cameroon and east to south¬ 
western Chad. Its distribution south of Cameroon and in East Africa is a matter of contention 
(Loveridge 1951; Rodel and Schmitz 2010). Current literature, including Hughes (1985), Chippaux 
(2006), Trape and Mane (2006) and Wallach et al. (2014), have not included Angola in the distri¬ 
bution of P. irregularis, although Wallach et al. (2014) included southern and western provinces of 
the Democratic Republic of Congo in its range. Based largely on the maps of Hughes (1985), we 
