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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
South Africa. Thelotornis capensis oatesi occupies much of this range exclusive of South Africa, 
eastern Botswana and adjacent areas. 
Ocurrences in Angola (Map 366): The 
subspecies occurs in southern and central-east 
Angola. Lunda Norte: “Sombo” [-8.68333, 
20.95000] (Laurent 1954:58). Lunda Sul: 
“Alto Chicapa” [-10.88333, 19.23333] (Lau¬ 
rent 1964a:116; Broadley 2001:68). Benguela: 
“Quisange” [-12.43333, 14.05000] (Bocage 
1895a:119; Loveridge 1944b:159); “Hanha” 
[-12.25000, 13.75000] (Bogert 1940:70; 
Loveridge 1944b: 159; Broadley 2001:68). 
HmTa: “Caconda” [-13.73333, 15.06667] 
(Loveridge 1936a:40, 1944b: 159; Broadley 
2001:68); “Quillengues” [-14.06667, 
15.08333] (Bocage 1895a: 119; Loveridge 
1944b:159); “Kuvangu” [-14.46667, 16.30000] 
(Monard 1937b:135; Loveridge 1944b:159); 
“Huilla” [-15.05000, 13.55000] (Bocage 
1895a: 119; Broadley 2001:68), “Joa Almeida” 
[-15.18333, 13.68333] (Broadley 2001:68). 
Namibe: “Biballa” [-14.76667, 13.36667] (Bocage 1895a:119; Loveridge 1944b:159). Cunene: 
“Chimporo” [-16.03333, 17.15000] (Monard 1937b:135; Loveridge 1944b:159); “Dorbush am 
Cunene, Chitado (thorny bush in Chitado)” [-17.30000, 13.90000] (Hellmich 1957b:69; Broadley 
2001:68). Cuando Cubango: “Cuito basin (52)” [-14.59333, q] (Conradie et al. 2016:9, 12, 19). 
Undetermined Locality: “Rio Quando” (Bocage 1895a: 119; Loveridge 1944b: 159). 
Taxonomic and distributional notes: Bocage (1895a) noted that some Angolan specimens in 
the Museu Bocage collections previously identified as Dryiophis Kirtlandii from “Quissange,” 
“Biballa,” “Rio Quando,” “Quillangues,” and “Huilla” resembled the individuals described by 
Gunther (1881) as Dryiophis Oatesi from “Matabeleland,” Zimbabwe. Loveridge (1944b) consid¬ 
ered two races of Thelotornis kirtlandii (Hallowell, 1844), one from the north as the typical form 
and one from the south as Thelotornis kirtlandii capensis, which might represent the southern 
records of Bocage (1895a) and Monard (1937b). This association should correspond to Thelotor¬ 
nis capensis oatesi as suggested by Loveridge (1953a) and represented in Broadley (1979). 
Broadley (1979) and Broadley and Wallach (2002) presented distribution maps but did not list the 
localities plotted. The status of T c. oatesi as a valid taxon remains problematic and has not been 
resolved by morphological and genetic analyses (Eimermacher 2012). 
I2"E 16.°E 20°E I4'E. 
Map 366. Distribution of Thelotornis capensis oatesi in 
Angola. 
Thelotornis kirtlandii (Hallowell, 1844) Forest Twig Snake 
Leptophis Kirtlandii Hallowell 1844:62. Holotype: ANSP 5271 (collector E. Blanding, don. W. Blanding). 
Type locality: “Liberia, West Africa.” 
Dryophis Kirtlandi: Bocage (1866a:48). 
Thelotornis Kirtlandi'. Ferreira (1900a:52), Boulenger (1915:213), Parker (1936:125), Themido (1941:10), 
Laurent (1954a:59, 1964a: 116), Thys van den Audenaerde (1966:34). 
Thelotornis kirtlandii eapensis: Loveridge (1944c: 154). 
Thelotornis kirtlandii kirtlandii'. Loveridge (1944c: 149, 1957:274), Hellmich (1957b:69). 
Thelotornis kirtlandii'. Spawls and Branch (1995:24), Broadley (2001:57), Broadley and Wallach (2002:69), 
