FAULL: DEVELOPMENT OF ASCUS. 
95 
11, fig. 76). Sometimes, however, the tip is segmented off in the 
conventional manner, and then the ascus grows out from the penul¬ 
timate cell. Not all the nuclear phenomena have yet been eluci¬ 
dated, but such as have been, present no new features. 
Early in the development of the ascus, extranuclear bodies make 
their appearance about the “fusion” nucleus. They are in the 
form of granules or Hat masses which lie close against the nuclear 
wall. They are not scattered about the protoplasm or away from 
the nucleus as was found to be the case in certain stages in the 
development of the asci of Hydnobolites. Figures 60 and 61 (pi. 
10) represent similar bodies in species of Podospora, in which, how¬ 
ever, they are larger than in Sordaria fimicola. The number of 
these bodies is at first small, perhaps not more than one or two, but 
later on there may be a dozen of them around a single nucleus. 
Just before the first spindle is formed they disappear and do not 
again occur. They are probably homologous with Guilliermond’s 
“ corpuscules metachromatiques.” 
There is the same differentiation of protoplasmic contents that we 
have already observed in Neotiella albocincta, and the first nucleus 
lies within the thick protoplasm. This nucleus, however, does not 
contain nearly as much chromatin as the corresponding one in 
Neotiella, though the achromatic network is well developed. The 
centrosomes throughout are flattened and disc-shaped in the rest¬ 
ing nuclei, but smaller in those that are dividing. 
The first spindle is prominent, though very slender, is intra¬ 
nuclear, and possesses long, fine astral rays which are demonstrated 
with some difficulty. Its long axis lies in the long axis of the ascus 
or very nearly so. In connection with the second set of spindles 
there are no new features, and the only point to be mentioned is the 
same extreme lengthening in the anaphase stages that has been 
noted elsewhere. The resting nuclei at the end of this division 
attain a fairly large size, and their centrosomes are particularly 
clear (pi. 10, fig. 53). 
During the four-nucleated stage, a differentiation of the thick 
protoplasm may begin, though the heaping up of the protoplasm 
about the nuclei becomes much more obvious during the spindle 
stage that follows. From figure 54 (pi. 10) it will be seen that the 
protoplasm in the immediate neighborhood of the nuclei is finely 
reticulated, while farther away vacuoles or coarser reticulations are 
