MUNSON: SPERMATOGENESIS OF PAPILIO. 
75 
tion which it actually does occupy in the normal spermatocyte (pi. 13, 
fig. 36,. c). 
That which gives these cells their pathological aspect is the com¬ 
pactness of the nucleus. I have already called attention to similar 
nuclei represented in plate 13, figure 33, where their pathological 
nature is very evident since these cells actually crumble to pieces. In 
those cells, too, the nuclei are compact, deeply staining bodies devoid 
of karyolymph. I assume that the chromatin, for some reason, has 
ceased to function, inasmuch as the karyolymph — which I have 
found in other cells to be a product of chromatin, i. e., nuclear activity 
— is no longer produced. I should hesitate to consider as patholog¬ 
ical, the somewhat similar nuclei represented in plate 13, figure 32, 
were it not that in most cases after mitosis, the daughter nuclei are 
soon re-formed and expanded. 
The large cells in plate 13, figure 33, with several nuclei suggest 
that while the nuclei have divided into two, four, or even six distinct 
nuclei, the cytoplasm has evidently failed to constrict. The general 
appearance of this cytoplasm is certainly pathological; but it is diffi¬ 
cult to decide whether the disorganization is due to the nucleus or to 
the cytoplasm. The evidence will hardly permit the conclusion that 
synapsis is a diseased condition leading up to the final disorganization 
represented in plate 13, figure 33. (Compare pi. 13, figs. 24, 25, 26, 
27, 28, 32, 33.) 
Period of growth .— It seems certain, judging from the gradual 
increase in size of the gonocysts, that there is a slight period of growth 
after each mitosis during the division period. But it is equally clear 
that there now intervenes a considerable rest period in which the 
spermatogone, now a spermatocyte, simply grows. I have sought to 
represent these facts in the diagram (text-fig. A). 
The size finally attained by the spermatocytes varies greatly. At 
maturation most spermatocytes are comparatively minute (pi. 13, 
figs. 29, 30, 31). These minute ones are usually found near the 
periphery, the cytocvsts being scattered among the gonocysts. Occa¬ 
sionally, in the same testis with the former, there are found, nearer 
the center of the follicle, and among the mature spermatocysts, enorm¬ 
ously developed cytocysts with unusually large spermatocytes. Hence 
there are ordinary spermatocytes and giant spermatocytes (pi. 13, 
figs. 35, 36, 37, 38; pi. 15, figs. 76, 78, 79, 80). 
Under the heading “Stalked cells,” and in connection with cyto- 
