312 Brooks and Stiles .— The S'truehire of 
one another and fuse by their margins to form a V-shaped bundle with its 
xylem and phloem inversely orientated with regard to the xylem and 
phloem of the subtending bract bundle (PI. XXI, Fig. 13 ). This V-shaped 
bundle next straightens itself out (PI. XXI, Fig. 14 ) and begins to divide into 
three (PI. XXI, Fig. 15 ), so that as the chalazal end of the ovule is approached 
there are found three bundles, each, as a rule, with a small non-functional 
resin canal (PI. XXI, Fig. 16 ). 
At the chalazal end of the ovule the two lateral bundles bend round 
through about a right angle and descend, one on each side of the ovule, for 
a short distance towards the micropyle; as seen in transverse section they 
lie on the diameter of the ovule (PL XXI, Fig. 17 ). The central bundle 
continues its course over the top of the ovule, and in so doing divides into 
two bundles (PL XXI, Fig. 18 ) which descend a short distance towards the 
micropyle (PL XXI, Fig. 17 ). All these descending bundles soon die out, 
and all show a tendency to amphivasal structure (PL XXI, Fig. 20 ). 
The nucellus of the ovule is surrounded by two integuments. The 
inner is fused for about half its length with the nucellus. The outer 
integument or epimatium is free from the inner for a short distance at the 
micropyle end, and on the side remote from the ovular stalk; on the other 
side it is either not present or is completely fused with the stalk . 1 
Where free, the innermost layer of the inner integument often con¬ 
sists largely of resin cells. These are apparently continued into the part of 
the ovule where the inner integument and nucellus become fused, and 
so apparently mark the line of division between these. A zone containing 
larger and more numerous resin cells also seems to mark the line of 
division between the inner and outer integuments, but these do not extend 
far enough towards the chalaza, and are not definite enough to enable one 
to state whether the descending bundles belong to any particular integument 
or to the nucellus. 
In a species of Podocarpus grown in the Cambridge Botanic Garden, 
the structure was found, on the whole, to be similar. The peduncle is borne 
in the axil of an ordinary foliage leaf. In the stage examined the peduncle 
is about six millimetres long and nearly a millimetre broad, and, as in 
P. spinulosus , is flattened in a vertical plane at right angles to that con¬ 
taining the peduncle and the branch bearing it. In the few specimens 
available for examination the lowest pair of leaf-like bracts were not 
present, and the whole fructification consisted of two pairs of bracts fused 
together, of which one or both of the lower pair bore ovules. The 
arrangement of the bundles is practically the same as in P. spinulosus. 
One bundle of the stem passes into the subtending leaf, while the two 
bundles on each side of it pass into the peduncle. These almost imme¬ 
diately divide up into a ring of bundles. At the base of the fused bracts 
1 See Postscript. 
