Podocarpus spinulosus, ( Smith) R. Br. 313 
two sets of three bundles are present; these belong to the fertile scales. 
One of these bundles on each side divides, and so gives rise to the bundles 
of the two sterile scales. Additional small bundles may be present on 
the inside of the sterile scale bundles as in P. spinulosus . The xylem 
of the bundles tends to be semicircular in shape, more or less enclosing 
the phloem on three sides. The resin canals, of which there is one on 
the phloem side of each bundle, are much larger than in P. spinulosus , 
and as a rule the resin duct divides after the division of the bundle to 
which it belongs. The ovular bundle, as in P. spinulosus , divides into 
three; when on a level with the nucellus, transfusion tissue begins to 
appear both centripetally and laterally, and the division into three bundles 
is no longer obvious. These bundles die out at the chalaza, and do not 
descend on the sides of the ovule as in P. spinulosus . For the last part 
of their course they are composed of transfusion-like tracheides only. 
In Podocarpus alpinus , material of which was obtained from the 
Royal Gardens, Kew, the peduncle is very short, being less than a milli¬ 
metre long. The lowest pair of bracts are extremely small and scale¬ 
like, although Pilger (13) states they are absent altogether; of the next pair 
only one bears an ovule, while of the upper sterile pair one is often aborted 
altogether, and the other is small compared with those of the lower pair. 
The vascular system at the base of the fused bracts consists of four 
bundles arranged along the sides of a square; of these one serves the 
fertile scale and two the ovule it subtends, the fourth serving the opposite 
sterile scale. The upper sterile scale has usually no vascular supply. 
The two ovular bundles fuse as in the other species described, but at 
first almost the whole of the xylems of the two bundles come in contact, so 
that the resulting bundle is almost concentric; further up, however, it 
straightens itself out. Nearer the chalaza it divides into two bundles which 
curve over the top of the ovule and descend a little way towards the micro- 
pyle on the sides of the ovule in much the same way as the lateral bundles 
do in P. spinulosus. In P. alpinus the resin canal accompanying each of 
these two bundles extends down to the level of the nucellus where it occurs 
on the inner margin of the outer integument, to which these bundles would 
appear to belong. Other canals also appear on the inner margin of the 
outer integument; these presumably correspond with the middle bundle of 
P. spinulosus, which bends over the ovule and then divides into two. On 
the whole, it would seem from their behaviour in P. alpinus that these 
bundles all belong to the outer integument, but the evidence at present is 
insufficient to decide this point. 
As far as the female fructification is concerned, the Podocarpeae 
obviously fall into two divisions; one in which the megasporophylls are 
aggregated into cones, and which comprises the monotypic genera Saxe- 
got haea and Microcachrys } the other in which the fertile megasporophylls 
